THE PHILIPPINE LAND MOLLUSKS COCHLOSTYLA 

 RUFOGASTER AND OBBA MARMORATA AND THEIR 

 RACES 



By Paul Bartsch 



Curator, Division of Mollusks and Cenozoic Invertebrates, United States 



National Museum 



COCHLOSTYLA RUFOGASTER AND ITS RACES 



A sending of a lot of specimens of what is herein described as 

 Cochlostyla rufogaster juani, by the Philippine Bureau of Science 

 for determination, has made it necessary to subject the entire complex 

 of Cochlostyla rufogaster, as well as some other species which have 

 sometimes been designated under this name, to a critical review, the 

 results of which are set forth in the following pages. 



All the specimens mentioned as having been collected by me were 

 obtained during the cruise of the United States Bureau of Fisheries 

 steamer Albatross in the Philippine Archipelago, 1907 and 1908. 



Lesson, in 1831, described, and figured on Plate 22 of his Illustra- 

 tions de Zoologie, a shell in the collection of the Duke of Rivoli. The 

 figure, I believe, is referable to the large conic race that inhabits the 

 region about Mariveles, Bataiin Province, Luzon. The shell figured 

 is a dead decorticated specimen, the loss of whose epidermis exposed 

 the red color of the later turns and the light peripheral zone. Lesson 

 states that the habitat from which the specimen came was unknown. 



Cochlostyla rufogaster as now conceived extends over central 

 Luzon from Benguet south to Mount Banahao and Mount Maquiling. 

 As is usual with Philippine land shells, we find that a certain degree 

 of differentiation has taken place in the shells in the various habitats 

 occupied, which makes it not only possible but desirable to recognize 

 certain zoogeographic races and to designate these as subspecies, 

 which is here done. 



The races of Cochlostyla rufogaster vary from elongate-conic to 

 ovate, from moderately slender to inflated. They range in height 

 from more than 90 mm to less than 50 mm. In shells with perfect 

 periostracum this is buff on the early whorls and wood brown on the 



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