314 PROCEEDINGS OF THE NATIONAL MUSEUM voi 



Subgenus Culicoides (Oecacta) Poey 



Oecacta Poey, 1851, IMem. Hist. Nat. Isla Cuba, vol. 1, p. 236. (Type: Oecacta 



furens Poey, monobasic.) 

 Psychophaena Philippi, 1865, Verh. Zool.-Bot. Ges. Wien, vol. 15, p. 628. — 



Wirth, 1955, Rev. Chilena Ent., vol. 4, p. 234. (Type: Culicoides venezuel- 



ensis Ortiz, as Psychophaena pictipennis Philippi, monobasic.) 

 Haematomyidium Goeldi, 1905, Mem. Mus. Goeldi, vol. 4, p. 137. (Type: 



Haematomyidium paraense Goeldi, monobasic.) 



Species with the second radial cell usually entirely dark; wing 

 usually with prominent pattern; male parameres separate; basistyle 

 with well-developed, foot-shaped or pointed, ventral root; ninth 

 tergum with well developed apicolateral processes; spermathecae 

 usually two, rarely one; hind tibial comb with four (rarely five or 

 seven) spines. 



This subgenus has undergone a tremendous development in the 

 Neotropical region, with the evolution of numerous groups of related 

 species. Groups of this subgenus found in Panama are daedalus, 

 copiosus (p. 332), iriartei (p. 342), scopus (p. 347), limai (p. 349), 

 acotylus (p. 362), reticulatus (p. 376), furens (p. 397), Jiuvialis (p. 407), 

 transferrans (p. 420), leoni (p. 426), dehilipalpis (p. 431), pachymerus 

 (p. 454), arubae (p. 464), and stigmalis (p. 466). 



The daedalus group 



This group consists of medium-sized species with very hairy wing ; 

 male genitalia with ventral root of basistyle slender, short and pointed, 

 paramere with the base not directed laterad, with a long anterior 

 process on the basal knob and with simple twisted tip and aedeagus 

 simple, V-shaped or Y-shaped and slender at the apex; wing with a 

 pale spot present straddling vein jVL, a pale spot lying adjacent to 

 the anterior side of vein Mj only in crescentis; female antenna with 

 sensoria present on segments in, xi-xv or on iii, v, vii, ix, and 

 xi-xv; female palpal pit broad and shallow or deep and opening by 

 a small pore. 



Panama species: 



The species of this group are most numerous in western Panama at 

 Almirante or in the Upper Tropical Zone at Cerro Campana and El 

 Valle, only a few species ranging to the Canal Zone. There are 

 seven or eight Panama species: antefurcatus, new species; commatis, 

 new species; crescentis, new species; daedalus Macfie; dunni, new 

 species; pampoikilus Macfie, and phaeonotus, new species. Culicoides 

 daedaloides, new species, is provisionally referred here but its position 

 is uncertain. 



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