BITING MIDGES — WIRTH AND BLANTON 363 



always present on some of the proximal flagellar segments, absent 

 on the last five except in venezuelensis; tibial comb with four or five 

 spines; two spermathecae present; ventral root of male basistyle 

 slender or foot-shaped; aedeagus often with stout basal arch and 

 modified tip with well developed lateral points; parameres with or 

 without ventral lobe or distal fringing spines. 



Panama species: Six: acotylus Lutz, carsiomelas Wirth and 

 Blanton, guyanensis Floch and Abonnenc, lanei Ortiz, venezuelensis 

 Ortiz and Mirsa, and wokei Fox. 



41. Culicoides wokei Fox 



Figure 44 



Culicoides wokei Fox, 1947, Kuba, vol. 3, p. 90 (female; Panama; fig. wing, palpus). 

 Culicoides aethionotus Wirth and Blanton, 1955, Bull. Brooklyn Ent. Soc, vol. 50, 



p. 121 (male, female; Panama; fig. wing, palpus, spermathecae, male 



genitalia). New synonymy. 



Female: Length of wing 0.78 (0.75-0.84, n=9) mm. 



Head. — Eyes narrowly separated, bare. Antenna with flagellar seg- 

 ments in proportion of 17:13:13:13:13:13:13:13:16:17:18:20:30, an- 

 tennal ratio 0.95 (0.93-0.96, n = 2); distal sensory tufts present 

 on segments iii to x. Palpal segments in proportion of 8:16:20:8:10, 

 third segment short and swollen, 1.6 (1.5-1.7, n = 8) times as long as 

 greatest breadth, with broad, shallow sensory pit. Mandible with 

 14 (13-15, n=7) teeth. 



Thorax. — Mesonotum dull, blackish brown on humeri, broadly on 

 each side of sensory pits and narrowly along sides to wing bases, disc 

 uniformly dark grayish to coppery pruinose. Scutellum uniformly 

 pruinose brown; postscutellum and pleuron blackish. Legs dark 

 brown, all femora pale at bases; fore and mid femora with subapical 

 and all tibiae with subbasal narrow pale rings; hind tibial comb 

 with four (n = 5) spines, the one nearest the spur longest. 



Wing. — Pattern as figured; pale spots whitish and extensive, in a 

 framework of narrow, connected dark linear markings, principally 

 along the veins; second radial cell dark; pale spot over r-m crossvein 

 very broad, a large pale area over base of wing, the dark area between 

 these pale areas quite short ; cell R5 with three pale spots, a small one 

 at end of second radial cell, an elongate one behind second radial cell 

 sometimes narrowly meeting the pale spot over crossvein and the 

 anterior poststigmatic pale spot, and the third pale spot extensive, 

 nearly filling apex of cell R5, broadly meeting anterior wing margin 

 and continuing along the margin as a narrow pale appendage nearly 

 to end of cell Mi; two elongate, pale, streak-like spots in cell Mi, the 

 distal one broadly meeting wing margin ; two pale spots in distal part 

 of cell M2, the distal one broadly meeting wing margin; pale spot in 



