Notes on Hawaiian Zonitidae and Succineidae 



By C. Montague; Cooke, Jr. 



INTRODUCTION 

 The material on which the present paper is based forms part of 

 the malacological collection of the Bishop Museum. In 1920 I 

 took to Philadelphia for comparative study the animals and shells 

 of Hawaiian representatives of the families of Zonitidae, Endodon- 

 tidae, and Succineidae and worked out there the anatomy of a 

 number of the Zonitidae and Succineidae, but was unable to com- 

 plete the analysis for lack of sufficient material. These notes deal 

 with several genera of Succineidae and with three of the rarest 

 genera of Zonitidae found in Hawaii. For comparison with the 

 Hawaiian forms the very rich collection of wet material in the 

 Academy of Natural Sciences was placed at my disposal in Phila- 

 delphia by Dr. Henry A. Pilsbry, to whom I am also indebted for 

 advice and many courtesies. Thanks are due also to Miss Helen 

 Winchester, who made the drawings for the two plates. 



ZONITIDAE 



GODWINIINAE 



Sykes^ proposed the generic title of Godwinia for Gould's 

 Vitrina caperata, basing his conclusions on the anatomical studies 

 of Godwin-Austen. From a further study of the anatomy it appears 

 that a new subfamily title is necessary. The shells are rather small, 

 8-13 mm. in diameter, of few rapidly enlarging whorls, and a large 

 aperture ; the embryonic whorls are more or less distinctly and 

 roughly, distantly and radiately costate ; the umbilicus is small and 

 circular. 



Unfortunately the animals in all the specimens examined were 

 much contracted. As shown by Godwin-Austen there are no shell- 

 lobes. Both right and left dorsal lobes are strongly developed. 

 The sexual orifice is situated rather high and is back of the pul- 

 monary orifice (fig. 3, a). There is no tail pore or slit. The penis 

 is entirely different from that of other zonitoid snails. It is rather 

 large with a very short stout retractor. The distal end is slightly 

 enlarged into a head and is very thick walled with a minute almost 



* Sykes, E. R., Mollusca : Fauna Hawaiiensis, ii, p. 277, 1900. 



[3] 



