S. H. SCUDDER ON PALAEOZOIC COCKROACHES. 105 



terminates a little beyond the basal third of the wing; the anal veins, six in number, are 

 simple and similarly arcuate. 



The wing is somewhat below the medium size, the fragment being 17.25 mm. long and 

 6.5 mm. broad ; probably the real length of the wing is 18.5 mm., and the breadth to the 

 length as 1 : 2.85. The upper surface of the wing appears to be exposed, and is that of 

 the left side. Germar speaks of the principal mediastinal and internomedian veins as 

 delicate. 



Germar confounded this species with that formerly described by him in Minister's BeitraVe 

 under the name of Bl. flabellata. It is indeed very close in general appearance, but if the 

 figure given in Minister's Beitrage is correct, two species belonging to different genera are 

 indicated. The principal difference is to be found in the upper half of the wing. In Etobl. 

 flabellata (as first described, and as we have restricted it), the mediastinal area is very nar- 

 row, and the vein terminates at about the middle of the costal margin ; in Gerabl. M"n- 

 steri, on the other hand, the area is very broad, and the vein terminates only just before 

 the apex. In Etobl. flabellata again the scapular area is extensive, and filled with many 

 veins ; in Gerabl. Miinsteri, the scapular vein is simply forked once. Or, to put it other- 

 wise, the branches of the basal half of the mediastinal vein of Gerabl. Miinsteri are trans- 

 ferred, in Etobl. flabellata, to another short principal vein, running above the mediastinal, 

 and which does not exist in Gerabl. Miinsteri ; while the scapular vein of the latter, amal- 

 gamated at base with the three-branched externomedian vein, is to be considered, in Etobl. 

 flabellata, as the basal branch of a four-branched externomedian vein. The close resem- 

 blance of the externomedian and internomedian areas in the two wings would have led 

 me to consider the illustration in Minister's Beitriige as simply faulty, were it not for the 

 following considerations : First, Germar makes no mention of any such error, but merely 

 quotes the reference in his synonymy. Second, there are several points of difference 

 besides those pointed out ; for instance, the shape of the wing, which is less tapering in 

 Etobl. flabellata, with a less arcuate costal, and a more arcuate inner margin ; the com- 

 pound branch of the internomedian vein, found just beyond the middle of the wing in 

 Gerabl. Miinsteri, is represented in Etobl. flabellata by a pair of forked branches, having 

 a widely distinct origin ; the simply forked vein which I have considered the scapular in 

 Gerabl. 3fi"uisteri originates from the externomedian vein much nearer the base than in 

 Etobl. flabellata ; and the borders of the broken tip do not agree in the two wings. Third, 

 if they are to be considered the same, the correct drawing is certainly the later one, but 

 the structure of the mediastinal vein is circumstantially described, as well as figured, in both 

 of Germar's works, in each case corresponding to the illustration in the same work ; yet the 

 structure of the wing of Etobl. flabellata is wholly in keeping with that of the genus Eto- 

 blattina, which comprises the largest proportion of the European palaeozoic cockroaches, 

 and is indeed very closely related indeed to that of Etobl. affinis and Etobl. anthracophila, 

 as we have already pointed out : and were it nut fur the remarkable similarity of the distri- 

 bution of the nervures referred in Etobl. flabellata to the externomedian vein, it scarcely 

 seems probable that any doubt would arise concerning the distinction of the two species. 

 Unless Germar's original types exist, and can be verified, it seems questionable whether the 

 point can really be decided. 



Germar, in his Wettin fossils, compares this species to Etoblaltina anaglyptica, which he 

 says it closely, resembles, so that one might take it for a small specimen of the same, but as 

 we have seen above, the Wettin species must be placed in Gerablattina and not in Etoblat- 



MRMOIKS BUST. sot'. NAT. HIsT. Vol.. III. 



