Proceedings of 

 the United States 



■IVT . 1 TIT \S^^ 



National Museum ^^^ 



SMITHSONIAN INSTITUTION • WASHINGTON, D.C. 



Volume 114 1963 Number 3467 



SCARAB BEETLES OF THE GENUS 



ONTHOPHAGUS LATREILLE NORTH OF MEXICO 



(COLEOPTERA: SCARABAEIDAE) 



Henry F. Howden^ and Oscar L. Cartwright' 



The purpose of this paper is to facilitate the identification of the 

 species of Onthophagus of the United States and Canada and to pre- 

 sent information on their habits and life histories. Twenty-three 

 species and subspecies described from North America, two species 

 introduced from Europe and Africa, and eleven previously unrecog- 

 nized species are included. A key to the species, bibliographical 

 references, complete new descriptions, photographs of both sexes, 

 and maps of distribution are given. Important nomenclatural 

 changes are presented. 



Acknowledgments 



As a staff member of the University of Tennessee, Howden was 

 encom-aged to make four field trips to Florida and Alabama to study 

 distribution and biology of the group. Later the Entomology 

 Research Institute of Canada supported three trips to Texas, Arizona, 

 and Mexico in 1958, 1959, and 1960. The Smithsonian Institution 

 sent Cartwi'ight, as a staff member of the United States National 

 Museum, to study types at the Museum of Comparative Zoology at 



J Entomology Research Institute, Canada Department of Agi-lculturc, Ottawa, Canada. 



* Division of Insects, United States National Museum, Smithsonian Institution, Washington, D.C. 



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