BEETLE GENUS ONTHOPHAGUS — HOWDEN, CARTWRIGHT 103 



male major but broader, with the basal portion of the four outer 

 teeth larger. 



Variation. — Length ranges from 5.0 to 7.1 mm. and the width 

 from 2.9 to 4.2 mm. Color of individuals varies from reddish brown 

 to completely black. The alutaceous sculpture, tubercles, and setae 

 show little variation except the slight sexual one noted above. The 

 greatest variation other than sexual differences is in size and shape 

 of the pronotal protuberance of the male which ranges from a rounded 

 hump in some male minors to the extensive projection described 

 for the male major. 



Type. — Canadian National Collection 7531. 



Type locality. — 1 mile east of Portal, Ariz. 



Specimens examined. — 151. 



Distribution. — (See fig. 7, p. 59.) Holotype and allotype, I mi. 

 east of Portal, Ariz., Jime 26, 1956, H. and A. Howden, in wood rat 

 nest (Neotoma sp.). And the following paratypes: 



United States: Arizona: 26 cfc?", 26 ? 9, 1 mi. east of Portal, June 26-29, 

 1956, II. and A. Howden, O. L. Cartwright, F. N. Young (cNC, usnm, Howden, 

 fny); 1 cf , 1 9 , Sept. 14, 1960, H. F. Howden, in Neotoma nest (cnc); 10 cf cf", 15 

 9 9, July 31, 1961, 4 9 9, Aug. 1, 1961, in Neotoma nest, L. J. Bottimer (ljb); 

 1 9 , Southwestern Research Station laboratory, 5 mi. west of Portal, at light, June 

 28, 1956, H. and A. Howden (Howden); 1 cf , 1 9 , Southwestern Research Station, 

 July 12, 1956, R. J. Frederick (rjf); 1 9, Southwestern Research Station, Aug. 

 28, 1958, elevation 5000-6000 ft., H. V. Weems, Jr. (rew); 1 9 , Southwestern Re- 

 search Station, July 15, 1961, B. Benesh (bb) ; 1 o", 1 9 • 1 mi. south of Portal, July 

 11, 1956, R. J. Frederick (rjf); 1 cf, 1 9 , Mt. Graham, July 28, 1954, at light, 

 F. G. Werner (ua); 1 9 , Mayer, July 1937, E. R. Leach (gas); 1 c^, Patagonia. 

 Santa Cruz Co., July 15, 1955, at light, F. G. Werner and G. D. Butler (ua); 

 I cf, Prescott, July 9, 1936, E. R. Leach (cas); 1 9, Mt. Graham, near Saflford, 

 July 14, 1956, Vincent D. Roth (rew); 1 d^, 1 9 , Chiricahua Mts., July 8, 1952, 

 R. H. Beamer (uka); 2 9 9, July 12, 22, 1953, D. J. and J. N. KnuU (osu); 1 9 , 

 Ramsey Canyon, July 10-15, 1941, A. Klots (amnh). new Mexico: 1 c?. State 

 label only, (H. W. Wenzel coll., osu). texas: 10 c?" cf, 14 9 9 , Chisos basin. Big 

 Bend National Pk., May 5, 19, 29, 1959, Howden and Becker, at black light and 

 on carrion (cNc). 



Mexico: chihuahua: 8 c? cf , 5 9 9 , Majalca Rd. (5500 ft.), 30 mi. northwest 

 of Chihuahua, April 17, 1961, wood rat nest, Howden and INIartin (cNC, Howden). 



Remarks. — This species can be readily separated from all other 

 United States Onthophagus except velutinus by the tuberculate pro- 

 notum and elytra and the dull black alutaceous sm-face between the 

 tubercles. The male majors are easily distinguished from velutinus 

 by the lack of horns on the head and the odd heart-shaped pronotal 

 projection. Male minors and females of browni are sometimes very 

 difficult to separate from females of velutinus, being separable only 

 by the shape of the carinae on the head and the larger pronotal hump 

 as noted in the descriptions and in the ke3^ It gives the authors 

 considerable pleasure to name this distinctive species in honor of W. J. 



