SOME MOTHS, GENUS ACLERIS — OBRAZTSOV 259 



Acalla macana [sic]. — Petersen, 1924, Lepidopteren-Fauna von Estland, p. 335. 

 Acleris maccana. — Obraztsov, 1955, Tijdschr. Ent., vol. 98, p. 193, fig. 357; 1956, 

 Tijdschr. Ent. vol. 99, p. 149. 



In his revision of tiie Canadian Acleris (=Peronea) species, Mc- 

 Dunnough (1934) treated Jishian a as a synonym of maccana, believing 

 that "there are a few very slight, minor differences between the geni- 

 taha of European and North American specmiens, but scarcely 

 sufficient to warrant the retention of the name jishiana Fern, even 

 in a varietal sense." In a more recent paper, McDunnough (1940) 

 changed his opinion, and appHed the name jishiana in a specific 

 sense. For support of his new point of view, he compared the geni- 

 talia of North American specimens oi jishiana mth those of the Euro- 

 pean maccana, and found some differences, especially in the structure 

 of the female genitalia. The most important difference consists of 

 the length of the antrum which m. jishiana extends over the cephalic 

 margin of the sterigma, being in maccana shorter and not reaching 

 this margin. Then, the caudal portion of the ductus bursae is in 

 jishiana somewhat expanded, "being largely membranous with only 

 indications of irregular chitinization," and not reaching the cephalic 

 margin of the sterigma. A further difference, found by AIcDunnough, 

 is the somewhat longer lateral lobes of the sterigma, which are in 

 Ushiana less outwardly oblique and have a small terminal spine. 

 In the male genitalia, he noted that in maccana "the apical section of 

 the tegumen shows a much narrower and shallower median excavation 

 and the lateral flaps are less evident," and "the apical recurvation of 

 the claspers is not so strong, resulting in a broader and shallower 

 excavation of tlie ventral margin than is the case in jishiana." Mc- 

 Dunnough emphasized that besides jishia7ia he also had from Canada 

 three female specimens of the true maccana which agreed with the 

 European specimens of this species. AlcDunnough was thus inclined 

 to believe that both jishiana and maccana are represented in the 

 Canadian fauna. 



Because of the variation of the genitaha in the European specimens 

 of maccana which show the same extremes as the characters described 

 by McDunnough as typical of jishiana, the present author cannot 

 subscribe to the opinion that jishiana and maccana are separate 

 species. Owing to the transitional shape of the genitalia in certain 

 specimens of maccana, it is impossible to recognize the above char- 

 acters as constant iov jishiana, the more so because even in the North 

 American specimens this transition has been reported by the present 

 author. A complete coincidence in the variation of the external 

 characters of European and North American specimens confirms 

 their conspecifity. The form jishiana corresponds to the nominate 

 form of maccana, and falls as its synonym. Besides this form, two 



