BREEDING TUBERCLES IN FISHES COLLETTE 569 



118 species of Percidae. I also examined nonbreeding material of all 

 species except Stizostedion marinum. 



Tubercle patterns, together with brief comments on other types of 

 sexual dimorphism, are described, species by species, in the following 

 sections. The suprageneric classification used follows that outlined 

 in a recent paper (Collette, 1963), where the Percidae was divided 

 into two subfamilies, the Luciopercinae and the Percinae. Each of 

 these subfamilies was subdivided into two tribes, the Luciopercini 

 and the Romanichthyini in the first subfamily, and the Percini and 

 Etheostomatini in the second. The arrangement of genera, subgenera, 

 and species within the Etheostomatini (table 1) is modified from that 

 presented by Bailey (in Bailey and Gosline, 1955). 



For the loan of specimens and other assistance, I wish to thank 

 Edward C. Kaney, Cornell University; Robert H. Gibbs, Jr., Ernest 

 A. Lachner, and W. Ralph Taylor, U.S. National Museum; Reeve M. 

 Bailey, University of Michigan; Royal D. Suttkus, TuJane University, 

 Frank B. Cross, University of Kansas; James E. Bohlke, Philadelphia 

 Academy of Natural Sciences; Clark Hubbs, University of Texas; Ota 

 Oliva, Charles University, Prague; GUes W. Mead, Museum of Com- 

 parative Zoology, Harvard University; Y. J. McGaha, University of 

 Mississippi; and Robert R. Ross, Virginia Polytechnic Institute. 

 Petru Banarescu of the Academia Republicii Populaie Romine 

 deserves extra thanks for sending specimens of all the Rumanian 

 percids to the museums of the United States. To Robert V. Miller, 

 Rudolph J. Miller, Leslie W. Knapp, and William J. Richards, I owe 

 a special debt of gratitude for their efforts in providing me with data 

 and material on percid breeding tubercles. Without the help of this 

 group of former feUow Cornell students, my work would have been 

 immeasurably more difficult. Mildred H. Carrington has drawn the 

 figures which valuably supplement this paper. Reeve M. Bailey, 

 Daniel M. Cohen, Robert H. Gibbs, Jr., Ernest A. Lachner, and 

 Rudolph J. Miller have all offered valuable suggestions on the 

 manuscript. 



Tribe Luciopercini 

 (Jenus Stizostedion Rafinesque 



There is little sexual dimorphism in the five species of Stizostedion — 

 canadense (Smith), lucioperca (Linnaeus), marinum (Linnaeus), vitreum 

 (MitchUl), and volgense (Gmelin) — and breeding tubercles are appar- 

 ently absent. Females of aU species reach a larger maximum size 

 than the males and on the average are longer and heavier in a given 

 year class, at least after the first two years of life. For S. canadense, 

 see Carlander (1950); for S. vitreum see Eschmeyer (1950), Hile (1954) 



