BREEDING TUBERCLES IN FISHES — COLLETTE 



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the second dorsal fin and on the anterior surface of the upper pectoral 

 rays. None are present on the first dorsal, anal, or pelvic fins. Two 

 other males with greatly enlarged testes (MCZ 3063) and 7 egg-filled 

 females (USNM, MCZ, ANSP) lack tubercles. Dr. MHler also 

 examined a large female Z. zingel in the Charles University collection 

 which had poorly developed tubercles on the head scales, scales of the 

 anterior dorsum, and on the pectoral rays. Vladykov (1931) reported 

 that sexual dimorphism was rather marked in this species, but only 

 mentioned that the paired fins of the males were much longer than 

 those of the females. 



I examined one male of Zingel asper with well-developed testes 

 (ANSP 14055, France, Rhone R., 116 mm.) and one female filled 

 with large eggs (MCZ 2551, Saone E,., 105 mm.). Neither had any 

 breeding tubercles. Both specimens, however, were in poor condition, 

 and Z. asper will probably be found to have a tubercle distribution 

 very similar to that of its close relative Z. streber. 



Genus Romanichthys Duniitrescu, Banarescu, and Stoica 



I have examined four tuberculate males of the only species in this 

 genus, Romanichthys valsanicola Duniitrescu, Banarescu, and Stoica 

 (USNM 190149, Arges R., May 22, 66 mm.; USNM 187749, Arges R., 

 May-June, 68 mm.; AICZ 40966, Vilsan R., June-Aug., 85 mm.; and 

 UMMZ uncat., Arges R., June-July, 76 mm.) and two tuberculate 



Figure 1. — Male of Romanichthys valsanicola showing the distribution of breeding tubercles 

 (USNM 190149, Rumania, Arges River, May 22, 66 mm. SL). 



females (MCZ 40966, 86 mm. and UMMZ uncat., 69 mm.). Another 

 male (USNM 190180, 92 mm., June-Aug.) has well-developed testes 

 but lacks tubercles. Four females (USNM 190149, 96 mm.; MCZ 

 40966, 80 and 89 mm.; and UMMZ uncat., Vilsan R., Oct. 19, 65 mm.) 

 also lack tubercles. The tubercles are best developed in the smaller 

 USNM male (fig. 1). There are a few small tubercles distally on the 

 dorsal surface of pelvic soft rays 2-5 and ventrally on rays 3-5. 



