600 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 117 



tubercles on the anal fin rays and on the belly scales of the paratj'-pes 

 described above. 



Males of E. imnctulatwm in seven collections (UMAIZ, KU, CU, 

 TU) taken from January 30 (TU 22993, Oklahoma, Neosho R.) to 

 May 2 (TU 2251, Oklahoma, Arkansas R.) have tubercles. They are 

 present on the anal spines and soft rays of males taken in this period 

 but are best developed on the soft rays. A 54 mm. male (CU 37346, 

 Missouri, Osage R.) taken March 28 has the maximum tubercle 

 development observed. There are 15-20 tubercles on each anal soft 

 ray, fewer on the anal spines. Tubercles are present on the ventral 

 surfaces of aU pelvic fin elements, ranging in number from 5 (on the 

 spine) to 30 (on the second soft ray). Tubercles are present on the 

 dorsal surfaces of pelvic soft rays 1-3 as well as on the ventral sm-faces. 

 Tubercles are also present on about 7 rows of the ventral scales per 

 side just anterior to the genital papilla and on about 6 rows along each 

 side of the anal fiii base. There are a few small tubercles on the ventral- 

 most principal caudal ray. 



Males of E. luteovinctum are tuberculate from at least March 26 

 (CU 37235) imtU April 28 (UMMZ 121612) in Duck River, Tennessee. 

 Tubercles develop earliest and remain for the longest period of time 

 on the anal fin. Except for tubercles on the anal spines, the males in 

 CU 37235 have the best developed tubercles of the specimens ex- 

 amined. No tubercles are present on the first anal spine, a few are 

 present on the second anal spine, and there are 5-10 tubercles on each 

 of the anal soft rays. There are no tubercles on the ventral surfaces 

 of the pelvic fin elements, but four intermediate-sized specimens have 

 several tubercles distally on the dorsal surface on soft rays 1 and 2. 

 Ventral tubercles start halfway from the pelvic to the anal fin and 

 extend onto 4-5 scale rows per side just anterior to the genital papilla, 

 3 rows along each side of the anal base, and 5 midventral rows on the 

 caudal peduncle. A few tubercles are present on the ventralmost 

 principal caudal ray in the largest specunen. No tubercles are present 

 on the pectoral fin. Some males (UMMZ 120985) have tubercles on 

 the first instead of the second anal spine. Others (UMMZ 121259) 

 have tubercles on both anal spines, although the tubercles are not as 

 well developed in the other regions as in CU 37235. The nine collec- 

 tions with tuberculate males contained no tuberculate females. 



Tubercles are present on the anal, pelvic, pectoral, and caudal fins 

 and on the ventral scales of the males of E. spectahile. I have found 

 tubercles on males of E. s. spectahile taken in late March (CU 37240, 

 Missouri, Osage R.; CU 37496, Tennessee, Barren R.; CU 37436, 

 Arkansas, White R.) and in early April (CU 38406, Missouri, Osage 

 R.). Tubercles are present on 61 males (26-42 mm. standard length) 



