580 PROCEEDINGS OF THE NATIONAL MUSEUM vol. u 



occasionally fused with adjoining intersegmental membrane showing 

 in this case a somewhat modified structure; lamella postvag- 

 inalis generally not defined, membranous, forming dorsal wall of 

 sinusal funnel. Antrum tubular, generally sclerotized, always 

 rather short and wide, as a rule with two lateral colliculi, one on each 

 side of antrum. Corpus bursae generally elongate, much longer than 

 broad, occasionally somewhat ovate, in greatest portion membranous 

 with very fine sculpture; cervix: bursae as a rule not separated from 

 corpus bursae, merely definable morphologically, more or less sclero- 

 tized and scidptured, as broad as or broader than antrum, occasionally 

 fused with it; ductus seminalis opening into caudal portion of cervix 

 bursae, close to antrum; cestum elongate, more or less sclerotized, 

 often with a longitudinal carina, located or starting in cervical area 

 of bursa copulatrix, rarely cephalad of it or missing; signum generally 

 present, shaped as slightly sclerotized, small area or plate, not always 

 distinctly separated from adjacent surface of corpus bursae, gen- 

 erally with a transverse, slightly elevated carina and/or various 

 sculptures. 



Remarks. — The valva of the Polyorthini is very peculiar and unique 

 for the entire family Tortricidae. If spread, it is extraordinarily 

 broad, but normally it is folded along the sacculus, and the broad 

 infrasaccular lobe, unbent externally, forms together with the inner 

 lobe of the valva a kind of pocket opened externally and costally. 

 This pocket serves as a receptacle for the cluster of long hairs of the 

 corema. The folded valva is foliaceus in its shape. Morphologically 

 the external lobe of the valva is probably the same as the infra- 

 saccular portion of the valva of some species of Archipini and Ana- 

 crusiini, in which it is rudimentary and not straight. 



The female genitalia of the Polyorthini are characterized by a wide 

 ostium bursae, and a low differentiation of the separate sections of the 

 bursa copulatrix. In most species of the tribe, merely the antrum is 

 sejDarated from the bursa while the cervix bursae is united with the 

 corpus bursae. Only some few species have the caudal portion of the 

 cervical section narrower than the remaining corpus bursae. The 

 ductus seminalis opens mto this narrow portion, but in the species 

 ^^•ithout this contraction the cervical section of the corpus bursae 

 directly receives the ductus seminalis. In connection with this mor- 

 phology, there are reasons to indicate the caudal sclerotized sculpture 

 of the wall of the corpus bursae as a cestum, leaving the name of the 

 signum for the sculpture located closer to the fundus bursae. 



In regard to the genitalia, the tribe Polyorthini is rather close to the 

 Cnephasiini. The position of vein R5 of the forewing, running in the 

 Polyorthini to the costa or the \nng apex, speaks rather for a more 

 ancient origin of this tribe and its probable relationship to the Tor- 



