NO. 3544 SUBSPECIES OF CAMBARUS LONGULUS — JAJMES 3 



placed longirostris with C. longulus as a subspecies of the latter (p. 

 966) and characterized both subspecies in a clear, concise key to the 

 North American Astacidae. 



1903. Harris, in a compilation of data (1903, p. 58), referred to 

 C. b. longulus and C. b. longirostris. I have presumed this to be in 

 error owing to Harris' interpretation of Faxon (1885a). 



1905-1912. Having erected four subgenera of Cambarus, Ortmann 

 established the subgenus Bartonius (1905, p. 120), which was declared 

 a synonym of Cambarus for reason of priority by Fowler, who listed, 

 in the place of Bartonius, the subgenus Cambarus (1912, pp. 340-341). 



1914. For some unexplained reason, Faxon reduced longulus to 

 subspecific rank calling it ^^ Cambarus bartonii longulus" and retained 

 longirostris as "C. bartonii longirostris." He completely omitted 

 Hay's designation and ^^Tote: "The character of the suborbital margin 

 of the carapace seems to be very constant A\dthin the limits of a good 

 subspecies, and it may prove to be the really diagnostic feature for 

 separating C. b. longulus and C. b. longirostris" (pp. 389, 424). Fifteen 

 years before. Hay (1899, p. 966) had utiUzed this identical diagnostic 

 feature in his key. 



1931. Ortmann, follo^\•ing Fowler, listed both "Cambarus 

 (Cambarus) longulus longulus Girard (1852)" and "Cambarus (Cam- 

 barus) longulus longirostris (Faxon) (1855)" (pp. 118, 121). Ortmann 

 considered the type locality for longirostris to be "The first exact 

 locality given by Faxon ('85b) . . . Doe River, Carter Co., 

 Tennessee" (p. 121). 



1942. Ten years later, in a revision of the genus Cambarus, Hobbs, 

 primarily adopting Ortmann's ideas of subgeneric relationships, 

 elevated the previously recognized subgenera to the rank of genus 

 within the new subfamily Cambarinae. 



1959-1961. Hobbs (1959, p. 896) indicated that there are two 

 subspecies of Cambarus longulus and subsequently indicated to the 

 present -WTiter that a third (form) probably should be recognized, 



Cambarus longulus Girard 



The species Cambarus longulus Girard (1852) belongs to the Bartoni 

 Section (Ortmann, 1931, p. 105) of Cambarus Erichsoi (1846) (as 

 redefined by Hobbs 1942d, p. 354) and, as herein recognized, consists 

 of three subspecies: Cambarus longulus longulus, C. longulus longi- 

 rostris Faxon, and C. longulus chasmodactylus, new subspecies. 



Geographical limits — The limits of the range of the species C. 

 longulus, although more specifically designated within the geographical 

 discussions of the respective subspecies are: North: represented 

 by C. I. longulus, in the upper James drainage in Greene County 

 (on the eastern side of the Blue Ridge) and Highland County (to the 



