NO. 3568 MELANOTUS — QUATE AND THOMPSON" 63 



1953. Siskiyou Co.: Yreka, May 10, 1932. Solano Co.: Green Valley Falls, 

 Apr. 26, 1941. Sonoma Co.: Santa Rosa, April 1942; Sobre Vista, May 24, 1910. 

 Stanislaus Co.: Westley, June 8, 1948. Tehama Co.: Red Bluff, Apr. 14, 1928, 

 May 11, 1949; Vina, May 11, 1920; Western, Oct. 30, 1920. Trinity Co.: Carr- 

 ville, May 17, 1934, May 30, 1934. Tulare Co.: Kaweah; Sequoia Nat. Pk., 

 June 13, 1929, May 25, 1930; Visalia; Wood Lake, Mar. 28-May 3, 1947. 

 Tuolumne Co.: Hardin Flat, May 29, 1955. Yolo Co.: Winters, Apr. 23, 1950. 

 Yuba Co.: Marysville, June 5, 1933. Idaho. Bannock Co.: Lava Hot Springs. 

 Canyon Co.: Parma, May 18, 1930. Lincoln Co.: Shoshone, June 10, 1938. 

 Montana. Lewis and Clark Co.: Wolf Creek, July 4, 1938. Nevada. No 

 other data. Oregon. Baker Co.: Richland, June 14, 1940. Benton Co.: Cor- 

 vallis, July 2, 1945. Douglas Co.: Drain, May 19, 1914. Harney Co.: June 22, 

 1912. Jackson Co.: McLeod St. Pk., May 22, 1960; Medford, June 12, 1915, 

 INLay 14, 1954; Talent, May 4, 1954. Lake Co.: Lakeview, May 27, 1957. 

 Yamhill Co.: June 1913, May 1934. Utah. Cache Co.: June 26, 1954; Lewis- 

 ton, JNIay; Bear River, Logan, 1925. Juab Co.: Eureka, July 2 (with /. longulus). 

 Salt Lake Co.: Apr. 14, 1909; Ft. Douglas, June 7, 1926. Tooele Co.: Stockton, 

 June 22, 1917. Utah Co.: Dividend, June 1921; Provo, June 2, 1939; Provo 

 Cnyn., May 21, 1913. Weber Co.: Roy, July 14, 1957. Washington. Kittitas 

 Co.: Yakima River, 8 mi. south of Ellensburg, July 22, 1959. Walla Walla 

 Co.: Walla Walla, June 1936. Whitman Co.: Pullman, May 13, 1930; Wawawai, 

 May 13, 1944. British Columbia. Oliver, May 22, 1924; Vernon, May 1, 

 1924; Victoria, June. 



The subspecies longulus and oregonensis are closely related forms, 

 allopatric in distribution (fig. 3). Longulus differs from oregonensis 

 rather constantly in having a smaller average size, being lighter in 

 color, the nasale better developed, the fovea a little deeper, the 

 antennae less sharply serrate, and the flagellar segments smaller. 

 These features are subject to some variation and a small percentage of 

 specimens possesses characters of the other subspecies. The genitalic 

 characters, on the other hand, seem entirely constant within the ranges 

 of the respective subspecies. Specimens have not been seen which are 

 intermediate between the two subspecies in genitaUc characters. A 

 specimen from northern Utah is intermediate in size and color but 

 oregonensis on genitalic characters. Perhaps these forms have reached 

 full species status and there is no interbreeding between populations. 

 This is improven, however, and we have seen few specimens from the 

 borders of the two forms' area. In oiu- opinion, the close relationship 

 and allopatric distribution is best shown by the nomenclatorial rank 

 of subspecies. 



Van Dyke (1932) recognized the affinity of longulus and oregonensis 

 and was the first to group them as subspecies of one species. At the 

 same time he recognized the thu'd subspecies, franciscanus . He 

 presents a scheme, that our observations do not support, in which the 

 smaller and lighter longulus of southern California gets larger and 

 darker in populations to the north in the SieiTa Nevada and the Great 

 Basin. The lowland populations in central California supposedly got 



