46 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 121 



Genus Demoleus Heller, 1865 



Caligus. — Otto, 1821, p. 15. [Refers to C. heptatus only.] 

 Binoculus. — Nordman, 1832, p. 32. [Refers to B. sexsetaceus only.] 

 Dinematura. — Burmeister, 1833, p. 331. [Refers to D. sexsetaceus only.] 

 Nogagus. — Gerstaecker, 1853, p. 63. [Refers to N. productus only.] 

 Demoleus. — Heller, 1868, p. 199. [Type-species: D. paradoxus.] 



Female. — Frontal plate distinctly separate. Thoracic segments 

 2-4: free. Dorsal thoracic plates on segment 4 only. Abdomen 1- 

 segmented and with a dorsal plate. Caudal rami large. Oral area 

 with adhesion pads associated with first and second antenna, maxilliped 

 and an additional pad between bases of the maxilhpeds. Oral 

 appendages of usual type. The second antenna with terminal hook- 

 like spine reduced. Penultimate segment of second maxilla with 

 large spine and patch of stout setules on distal corner. Legs 1-4 

 biramose; all rami 2-segmented. Legs 5 and 6 present. Egg strings 

 folded. 



Male. — Body fonn of usual type. No dorsal plates present. 

 Oral area similar to that of female. Appendages similar to female 

 except for a modification on endopod of third leg of male. IVIales 

 of this genus are poorly known. 



Discussion. — At present there are 2 species in this genus. D. 

 heptatus, female, is well known and is amply recorded in the literature 

 principally as an external parasite of Hexanchus. D. latus is reported 

 by Shiino (1954) from Acanthidium eglantina (Jordan and Snyder) 

 and I have examined material from Squalus acutipinnis Regan. Both 

 of these hosts are members of the family Squalidae. 



The adhesion process between bases of maxillipeds is not found in 

 any other genus in this family. This character alone sets it apart 

 from other pandarid copepods. 



Key to Females of Genus Demoleus 



Genital segment covering the dorsal abdominal plate. Egg strings exposed. 



heptatus 



Genital segment only covering about }^ of the dorsal abdominal plate. Egg 



strings hidden latus 



Demoleus heptatus (Otto, 1821) 



Figures 218-237 



Caligus heptatus. — Otto, 1821, p. 15. 

 Caligus paradoxus Otto, 1828, p. 352. 

 Binoculus sexsetaceus Nordman, 1832, p. 32. 

 Dinematura sexsetaceus. — Burmeister, 1833, p. 331. 

 Nogagus productus Gerstaecker, 1853, p. 64. 



