NO. 3570 PANDARIDAE — CRESSET 53 



spinules. Outer seta not greatly displaced from other 3, as in female. 

 Leg 6 (fig. 267) located internal to distal corner of genital segment and 

 consisting of a small process bearing short setae. 



Discussion. — The genus Pagina is closely related to Dinemoura 

 Latreille, 1829, of which there are 4 known species. These 2 genera 

 have the following characteristics of the female in common: a wing- 

 lilie dorsal plate on the fourth thoracic segment; a 2-segmented abdo- 

 men, each abdominal segment bearing a dorsal plate, the first of which 

 is bilobed and the second single-lobed, and legs 1-3 similar and rela- 

 tively unmodified. Pagina can be separated from Dinemoura by the 

 fact that in Dinemoura the fourth leg is broad and conspicuously 

 lamelliform, whereas the fourth leg of Pagina is unmodified. 



Pagina is unhke all other known genera of this family in having 

 the rami of legs 2-4 3-segmented. 



Genus Echthrogaleus Steenstrup and Liitken, 1861 



Dmemowra.— Guerin-Meneville, 1837, pi. 35. [Refers to D. alata only.] 



Dinematura. — Dana, 1852, p. 60. [Refers to D. braccata only.] 



Echthrogaleus Steenstrup and Liitken, 1861, p. 380. [Type-species: E. coleop- 



tratus.] 

 Pandarus.— Thomson, 1889, p. 363. [Refers to P. armatus only.] 



Female. — Frontal plate distinctly separate. First thoracic segment 

 fused with cephalon. Dorsal thoracic plates on segment 4. Abdomen 

 1 -segmented. Abdomen concealed beneath genital segment. Caudal 

 rami joined to abdomen terminally. Oral adhesion pads present but 

 somewhat reduced. First antenna 2-segmented. Legs 1-4 biramose. 

 Leg 4 lamelliform. Legs 5 and 6 present. Egg strings long and 

 straight. 



Male. — No dorsal thoracic plates present. Abdomen 2-segmented 

 Legs 1-4 biramose. Leg 3 with modification on endopod. Leg 4 

 not lamelliform. Legs 5 and 6 present and not as reduced as in female. 

 Other oral and thoracic appendages as in the female. 



Discussion. — This genus is cosmopolitan as a parasite on the body 

 surface of elasmobranch fishes. E. coleoptratus and denticulatus may 

 be restricted to larger pelagic sharks whereas torpedinis has been re- 

 ported only from the ray, Torpedo occidentalis. The copepod is para- 

 sitic on the body surface of the host. 



This genus is closely related to Dinemoura but differs from it princi- 

 pally in having a 1-segmented abdomen. The males of this genus 

 have not been well described. The descriptions of coleoptratus and 

 denticulatus males have been amplified here. 



In 1899 Thomson described a copepod under the name of Dinematura 

 hamiltoni. This is obviously a member of the genus Echthrogaleus, 

 but the figures and description are too poor to assign it to any species. 



