1.7-2.6 X 1.3-1.6 mm, teeth acute, 0.3-0.8 mm long. Staminate 

 plants: corolla 2.1-3.2 X 0.9-1.5 mm, abaxial lobe 0.7-1.7 mm 

 long, other lobes 0.4-0.6 mm long, pilose patch between abaxial 

 filaments (sparse to absent); filaments 0.4-0.9 mm long, anthers 

 0.3-0.4 mm long; ovaries 0.2-0.4 mm long, style 1.3-1.7 mm long, 

 stigmas ca. 0.2 mm long; nutlets at most 0.2-0.4 mm long; calyx 

 3.4-4.5 mm long. Pistillate plants: corollas 1 .7-2.7 X 0.8-1 . 1 mm, 

 abaxial lobe 0.5-0.9 mm long, other lobes 0.3-0.4 mm long, pilose 

 patch between abaxial filaments (sparse to absent); filaments ca. 

 0.2 mm long, anthers ca. 0.2 mm long; ovaries 0.3-0.4 mm long, 

 style 1 .3-2.2 mm long, stigmas ca. 0.2 mm long. Fruiting calyces 

 (Figure 1 C) yellowish green to brown and slightly purplish 

 tinged, subbilabiate, tube constricted distally, adaxially 3.4-4.8 

 mm long, abaxially somewhat shorter, tube 1.7-2.2 mm wide, 

 teeth acute, the adaxial 0.3-0.6(0.9) mm and the abaxial 0.4-0.9 

 mm long; veins 5 (to 7). Nutlets maturing to 1.8 mm long. 



Distribution and Ecology. Southern Ecuador (Figure 1 D); 

 dry terrain alongside roads and in disturbed secondary vegeta- 

 tion; 1700-2300 m; flowering from May through July. 



Representative Specimens. Ecuador. Loja: near Vilcabamba, 

 2300 m, Hart 774 (GH); 1760 m, Hart 1276 (GH); near Vilca- 

 bamba, 1 700 m, Hart 1983 (GH) Holotype!; near Quilanga, 2280 

 m, Hart 1358 (GH). 



Lepechinia dioica is a distinctive species recognized by its 

 leaves which are lanceolate to ovate-lanceolate, have obtuse and 

 long decurrent bases, and are densely and minutely pubescent on 

 the adaxial surfaces; highly branched inflorescences with 5 to 8 

 secondary branches and caducous floral leaves; small fruiting 

 calyces whose tubes are constricted distally; very small teeth 

 0.4-0.9 mm long, and 5 (to 7) veins; the tiny corollas 2. 1 -2.7 mm 

 long; and dioecious breeding system. The epithet "dioica "alludes 

 to the breeding system. 



Lepechinia dioica is most closely related to L. mutica and L. 

 mollis, and shares with them densely tomentose, ovate leaves with 

 acute bases. It differs from them by its decurrent leaf bases; 

 distally constricted, slightly purplish, much fewer 5- (to 7-) 

 veined mature calyces; and generally more highly branched 

 inflorescences. 



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