in 1960, when I proposed the five subfamilies or distinct phyletic 

 lines in the Orchidaceae. These phyletic lines were shown to have 



manner 



om 



morphic features. In that phyletic spectrum Apostasioideae, 

 Cypripedioideae and Neottioideae occupy the lesser evolved or 

 primitive end, while Orchidoideae and Epidendroideae were 

 shown to be advanced or derived. With the new data on Palmor- 



subfamily 



common 



these subfamilies. The plants of Apostasia/Neuwiedia, Sele- 

 nipedium and Palmorchis are all terrestrial with fibrous roots, 

 plicate leaves, and have three-locular ovaries and apterous, scle- 

 rotic seeds. Moreover, Palmorchis shares with Vanilla, in addi- 



lm 



mbe 



rostellum and the prominent stigma. Neobartlettia, commonly 

 included in Palmorchis, needs to be reinstated because of the 

 lack of fusion between the column and the lip. These are the only 

 genera which I now regard to comprise the subtribe Vanillinae 



must be mentioned 



Dietsch 



sympod 



growth habit of the plants, their undifferentiated leaves and 



segments 



me 



tion and illustration, needs further elucidations. The presence of 



mussen 



is indeed unique in the genus, but I consider that this structure 



more 



Because of the presence of the calyculus in the plants of Leca- 

 norchis, this latter genus has always been assigned to the sub- 

 tribe vanillinae notwithstanding the very disharmonious 

 colunmar structures and the tunicate seeds. 



The remaining 



mentioned by 



Lindley and commonly followed even today, i.e., Epistephium, 

 Galeola and also Eriaxis, I consider to form a new subtribe, 

 galeolinae beside vanillinae. The plants of these genera 



227 



