Battandier and Trabut, and Mordvinkina (1936) state 
that only the lower floret of 4. pilosa is articulated, while 
Post (1983) says that none of the florets are jointed to 
the axis. Haussknecht (1894), according to Thellung 
(1911), noted the same variation and believed this species 
to be a variety of A. clauda, differing only in the man- 
ner of floret attachment. Mal’tsev treats A. pilosa as 
closely related to and derived from A. clauda by a side 
branch of the main phylogenetic trunk. Both species, 
according to him, have unequal glumes, a linear callus 
about 8 mm. long and a linear-elliptical articulation-sear. 
3. A.longiglumis Dur. A wild oat found in southern 
Spain (Willkomm and Lange, 1870), Algeria (Battandier 
and Trabut, 1904), Palestine (Post, 19383) and Greece 
(DeHalacsy, 1904). Mal’tsev believes it to have given 
rise to A. ventricosa on the one hand, and to A. strigosa 
sens. ampl. on the other. 
4. A. ventricosa Bal. A wild oat restricted to west- 
ern Algeria (Battandier and Trabut, 1904; Mal’tsev, 
1929). Mal’tsev derived this species from the preceding 
by a side branch of the main phylogenetic trunk. It re- 
sembles 4. longiglumis in having equal glumes and a long 
callus with a narrow linear scar. It differs in having only 
the lower floret articulated, as in A. pilosa. 
5. A.Bruhnsiana Grun. A wild oat restricted to the 
Apsheron Peninsulain eastern Transcaucasia, and treated 
by Mal’tsev as a subspecies of 4. ventricosa from which 
it differs by the larger size of its spikelet-parts. Although 
the two are now separated by some 2500 miles, we do not 
know what migrations they may have undergone in the 
past. The wide gap is partially spanned by A. longiglu- 
mis from which they may have evolved independently. 
L271. | 
