198 THE WINGS OF NEUROPTERA 



During the past year Dr. Needham and I have reviewed our study of 

 the wings of ni}Tmeleonid pupae by the examination of fresh material and 

 Figure 193 is based on one of our recent photomicrographs. 



In no pupal myrmeleonid wing examined by us, and we have examined 

 many, were we able to detect the costal trachea. The subcostal and radial 

 trachege were prominent in all, so too were the accessory tracheae of the 

 subcosta. In some of the forming cross-veins there were distinct tracheae 

 but in most cases the tracheae in the cross-veins were so indistinct that we 

 are not warranted in representing them in the figure. 



Returning to a study of the medial trachea, it can be seen by an exami- 

 nation of Figure 193 that this trachea is two-branched; that trachea Mi +2 

 is a direct continuation of the main stem of this trachea and that trachea 

 M3+4 first traverses the oblique vein and then after making a sharp bend 

 extends lengthwise the wing in line with the basal part of trachea Cui. 

 From this it can be seen that the vein which in the adult wing appears to 



Fig. 193. — Tracheation of a fore wing of a pupa of a myrmeleonid. 



be vein Cui is really a serial vein consisting in part of vein Cui and in part 

 of vein Ms +4; to this serial vein Tillyard has appHed the designation 

 cubitomedian and makes use of the notation Cui + M2. I see no objection 

 to the application of the term cubitomedian to this vein; but as the nota- 

 tion Cui -f M2 indicates an ordinary coalescence of veins side by side and 

 not the formation of a serial vein, I regard the notation Cui & M3+4 as 

 more appropriate. 



It will be noted that Tillyard designates the branch of media that 

 unites with vein Cui in this case as Mo. This is an unfortunate failure to 

 apply the uniform terminology; and there are several other authors who 

 are guilty of the same error in similar cases. The two branches of media 

 formed by the first forking of this vein are veins M1+2 and M3+4; it is 

 obvious that these branches are homologous with those thus designated 

 when media preserves its typical four-branched form. If one designates the 

 two branches of media as veins Mi and M2 the question what has become 

 of veins M3 and M4 is suggested. 



In the wing represented by Figure 193 the trachea M3+4 and Cui were 

 distinct although they came very near together at the end of the oblique 

 vein and the veins formed about them anastomose at this point. In the 



