spores represents several sporangia ‘more or less com- 
pletely immersed in the tissue’. He discusses, however, 
a second possibility, namely that the segments themselves 
are enormous sporangia united at the base. This interpre- 
tation he rejects because of the great size of the segments 
and especially because of the vascular strands which are 
stated to run through them. In the light of the informa- 
tion which has now been gained regarding the structure 
of Whittleseya, Goldenbergia and Aulacotheca, 1 believe 
that this alternative interpretation is more probably the 
right one. The unusual size and shape of the spores and 
their aggregation in very long and narrow groups which 
form a uniseriate whorl round a central cavity are fea- 
tures characteristic of the three genera named, and if the 
sporangia, as believed by Sellards, are really immersed 
in the tissue, this would be yet another point of agree- 
ment.’ Whittleseya is a misunderstood form despite the 
lucid description given by Halle (7). 
Recently Wodehouse (17) has revived the obsolete 
and erroneous opinion that Whittleseya is a ginkgoalean. 
Whittleseya elegans Newberry is the staminate fructifi- 
cation of Alethopteris grandifolia Newberry, (12) which 
Arnold has shown to bear T'rigonocarpus-like seeds. 
Whittleseya is a pteridosperm of the order Medullosae. 
The spores of Codonotheca are very large, elongate- 
elliptical bodies 0.28—0.32 mm. long and 0.18—0.20 mm. 
wide. They retain a yellow resinous color sometimes ob- 
secured by reddish-brown stain. The spores usually contain 
two rather large dense structures which may represent 
nuclei. There are also present smaller irregular masses of 
a black substance which Sellards regards as the remnants 
of the original food supply. The pollen-grains of Codo- 
notheca caduca resemble those of Dolerotheca fertilis (7) 
in size, shape and ornamentation. 
Sellards described (p. 89) the spores of Codonotheca 
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