whorls of foliar appendages. It has not been possible to 
find traces of internal structure by means of peel and 
maceration techniques. Material for anatomical study is 
extremely limited since only one specimen of this plant 
is known. It has been impossible, even under relatively 
high magnifications, to find a definite ridge which might 
have been a vascular strand. The aspect of the entire 
plant, however, suggests, by comparison with other 
known vascular plant-fossils of similar nature, that it 
must have been vascular in structure. 
An important aspect of Sphondylophyton hyenioides 
is the presence of leaf-like appendages or foliar organs in 
whorls. The verticillation is more regular than in Hyenia 
sphenophylloides Nathorst (7,12) or than in Hyenia ele¬ 
gant Krausel & Weyland (9,10) of the Middle Devonian 
of Norway, Germany, and other parts of Europe. The 
whorls in Sphondylophyton are regularly spaced (Fig. 
II), on the lower fragments of axes averaging four to six 
millimeters apart; on the several shorter pieces, the 
whorls are placed about one to three millimeters apart. 
The axes bearing these crowded whorls are further dis¬ 
tinguished by having the foliar organs curled into a bud¬ 
like mass. Consequently it is probably safe to assume 
that these portions are terminal. 
The form of the foliar organs is also of extreme in¬ 
terest. They were apparently flat, somewhat coriaceous, 
and possibly more or less stiff. Each appendage of the 
whorl is basally dichotomous into two equal or subequal 
truncate-linear segments. The “terminal” axes, how¬ 
ever, possess foliar organs of a different shape. The seg¬ 
ments of the appendages are slightly incurved, pointed 
and curled together into bud-like masses. Whether or 
not any significance attaches to this difference is stil a 
question. , 
That the lower portion of the axes was probably naked 
[27] 
