Axis non-articulated. Sporangia with a tendency to serial 
and collateral arrangement, sitting on the adaxial side of 
the sporophylls. Sporophylls composing cone-like fruc- 
tifications. *’ 
In a later paper’, Nemejc described the typical heter- 
osporous pteridophytic spores of Noeggerathia foliosa 
Sternberg. 
Modern plant morphologists recognize three phyletic 
lines among the vascular plants: Lycopsida, Sphenopsida, 
and Pteropsida, which presumably trace their common 
origin to the Devonian Psilophytales or Psilopsida. Jef- 
frey’ more than thirty years ago proposed two groups, 
Lycopsida and Pteropsida. The Lycopsida are micro- 
phyllous with adaxial sporangia and the Pteropsida are 
megaphyllous with abaxial sporangia. Scott’ distinguished 
two groups of microphyllous plants, the Lycopsida and 
Sphenopsida. The Lycopsida have spirally arranged 
leaves and non-articulated axes, while the Sphenopsida 
have articulated axes and whorled leaves. The Sphenop- 
sida include chiefly, the Equisetales, the Calamitales, and 
the Sphenophyllales. 
Fructifications of three species of Noeggerathia and 
one attributed to Tingia ( Tingiostrobus tetralocularis 
Kon’no) have been described. 
Lady Isabel Browne has considered this problem from 
its broader point of view. She concluded that these fruc- 
tifications belonging to the Noeggerathiales find their 
closest affinity with the Sphenophyllales. Kon’no’® con- 
sidered that they bear closer relationship to the Lycopo- 
diaceae, while Nemejc insisted that they are quite dis- 
tinct. Lady Browne compared the sporangia of the 
Noeggerathiales with those of Sphenophyllum dawsoni 
Williamson. In short, she regarded the Noeggerathiales 
as having simple pseudo-macrophyllous leaves, non-artic- 
ulated axes, phyllotaxy non-verticillate, and the fructi- 
[175 ] 
