THE ANATOMY OF BIRDS. — NEUROLOGY. 193 



directly connected with the skull — excepting when, as in a woodpecker, elongated branchial 

 elements of the hyoid bone take on such office by curling over the cranium (figs. 73, 74). 



Section of the bone is required for further examination of the ear-parts. On longitudinally 

 bisecting the skull, or otherwise gaining access to the brain-cavity, the internal surface of the 

 periotic bone is brought into view (fig. 70, po, op, ep). It is the same bone we have seen in 

 the tympanic cavity, now viewed upon its cerebral surface. In a skull of any size, as that of the 

 eagle before me (from which the rest of my description will be taken), there is no difficulty in 

 making out the parts, although the periphery of the periotic bone is completely consolidated 

 with its surroundings. The periotic, or petrosal (Lat. petrosus, stony — from its hardness), or 

 " petrous part of the temporal," is the bony capsule of the inner ear, enclosing the labyrinth or 

 essential organ of hearing, — in fact, it is the skull of the ear, sometimes therefore called the 

 otocrane — just as ethmoidal parts form the "skull of the nose," and the sclerotal bones represent 

 a " skull of the eye." The periotic consists of the three bones already often mentioned, — the 

 prootic, po, epiotic, ep, and opisthotic, op, or anterior, superior, and posterior otocranial bones, 

 completely consolidated together, as well as with surrounding bones. The petrosal appears as 

 an irreg-ilar i)rotuberance in the inner wall of the brain-cavity, at the lower back part. It 

 seems to be more extensive than it really is, because the great superior semicircular canal, too 

 large to be entirely accommodated in the petrosal, has invaded the occipital bone, — the track of 

 its bed in that bone being sculptured in bas-rehef (fig. 70, asc). Behind this semicircular trace, 

 the deep groove of a venous sinus is engraved in the bone, making the tract of the canal still 

 more prominent (fig. 70, sc). The top of the petrosal and contiguous occipital is the floor of 

 a recess or fossa in which is lodged the great optic lobe of the brain, partly divided from the 

 general cavity for tlie cerebral hemisphere by a bony tentorium, like that which in mammals 

 separates the cerebellar from the cerebral fossee. On the vertical face of the petrosal, or on the 

 corresponding occipital surface, is a large smooth-lipped orifice, at least ^^ of an inch in longest 

 diameter ; it leads to a tongue-like excavation of the bone, in which the flocculus of the cerebel- 

 lum is lodged. In front, between the petrosal and alisphenoid (or in the conjoined border of 

 one or the other of these bones) is a considerable foramen, conducting the second and third 

 divisions of cranial nerve 5 (see p. 183; figs. 70, 71, ^) into the orbit. Beh)W the petrosal (in 

 fact, between the opisthotic and the exoccipital), near the border of the foramen magnum, is a 

 foramen (which may be subdivided into foramina), representing the foramen lacerum posterius 

 of mammals, transmitting cranial nerves 9, 10, 11 (see p. 183; fig. 70, 8). The general space 

 under description is continued to the margin of the foramen magnum by the exoccipital (fig. 

 7U, eo). Now on the vertical face of the petrosal itself — behind foramen for 5, above that for 

 9, 10, 11, in front of the large floccular orifice, will be seen a smooth-lipped depression, tlie 

 meatus auditorius internus (fig. 70, 7), at the bottom of which are at least two separate small 

 foramina. A bristle passed in the upper (or anterior) one of these two holes emerges outside 

 the skull, in the tympanic cavity, near the tympanic end of the eustachian tube ; it has traversed 

 the interior of the petrosal, in a track known as the fallopian nerviduct; it transmits cranial 

 nerve 7 — the facial, or portio dura. A bristle passed into the other of the two foramina may 

 also be made to come out in the tympanic cavity, but by a different track, for it emerges through 

 either the fenestra ovalis or the fenestra rotunda ; it has traced the course of cranial nerve 8, — 

 the auditory nerve or portio mollis. Both bristles have entered the common internal auditory 

 meatus, but the second one has traversed the ear-cavity proper, through the labyrinth of the 

 ear, and come out at the tympanic vestibular orifim (fenestra ovalis), or at tlie tympanic cochlear 

 orifice (fenestra rotunda). Either passage is easily made, without breaking down or indeed 

 meeting with any Ixmy obstacle, which would not be the case with a mammal. Cranial nerves 

 7 and 8 w(!re formerly comited as one (seventh) ; hence the name portio dura (" hard portion") 

 for the former, and 2)ortio mollis (" soft iiortion ") for the latter. The former, as said, traverses 

 the petrosal bone and escapes upon the face ; the latter, which is the true acoustic nerve, or 



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