Systematics 

 Acanthonotozomatidae 



A single pattern of sculpturing and body ornamentation is shared 

 by almost all species of Acanthonotozomatidae. Distinctions among 

 the 21 genera are found primarily in the mouthparts. That so many 

 genera and their species could be based on a differentiation of mouth- 

 parts and gnathopods while retaining a common pattern of dorsal 

 teeth and epimeral structure induces concern over the generic validity 

 of the buccal distinctions. Only the genus Odius Liljeborg has a 

 simplified sculpture and unmodified coxae. Specialization of the 

 mouthpart bundle for piercing and sucking may have resulted in 

 a loss of function of various parts not now under conservative 

 genetic control. Intergeneric differences in mouthparts may therefore 

 be non-conservative and only of specific value. 



The species to be described below illustrates this concern. According 

 to Hurley's (1954b) key, the new taxon should be assigned to Pariphi- 

 media Chevreux (1906). Its first maxillary palp is short and uniarticu- 

 late. The new species differs from the type species of Pariphimedia 

 by the incised lobes of the lower lip but in all other characters it 

 fits the diagnosis of Panoploea Thomson. To create a new genus 

 for a species sharing generic characters of Panoploea and Pariphimedia 

 seems unwarranted, especially in view of the fact that the two genera 

 differ by only two pairs of alternatives. Hence, the species is assigned 

 to Panoploea in the belief that the first maxillary palp character 

 represents a degenerative specialization whereas the lower lip character 

 represents a primordium. 



Panoploea(?) hedgpethi, new species 



Figures 1-2 



Diagnosis. — Body of the "normal" form, with two dorsal teeth 

 on each of pereonites 7, pleonites 1, 2, and 3; all pleonal epimera 

 with one posteroventral tooth, third pleonal epimeron also with 

 one upturned, posteromedial tooth slightly serrate ventrally, ventral 

 tooth slightly serrate ventrally only on third pleonal epimeron, 

 second and third epimera with posteromedial margins having faint 

 suggestions of one rudimentary tooth; lateral view of pereonites 



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