GAMMARIDEAN AMPHIPODA 139 



Hyale rubra frequens (Stout), new combination 



Hyale rubra (Thomson) : Hurley, 1957, pp. 910-913, figs. 30-50 (with synonymy). 

 Allorchestes frequens Stout, 1913, pp. 650-651. 



Hyale nigra (Haswell) : J. L. Barnard, 1962c, pp. 153-156, figs. 19, 20 (not Has- 

 well, 1880 nor other references to H. nigra and H. niger). 



No definite assignment of H. nigra to H. rubra is made except 

 for the H. frequens component listed by Barnard (1962c), although 

 H. nigra as shown sketchily by Schellenberg (1928), by Haswell 

 (1880), and described by Stebbing (1906) probably represent H. 

 rubra also. 



Diagnosis. — Differing from the New Zealand subspecies as re- 

 viewed by Hurley (1957) in the following characters: dactyli of 

 pereopods 1-5 very minutely serrated; only the distalmost spine 

 of article 6 of pereopods 1-5 very minutely striated; article 2 of 

 male gnathopod 2 stouter distally; posterior setae of article 6 on 

 female gnathopods 1-2 and male gnathopod 1 less abundant and 

 more sharply confined to a narrow space; distal setae of article 4 

 of maxillipedal palp shorter; ramus of uropod 3 very slightly longer. 



Barnard (1962c) synonymized Hyale frequens (Stout, 1913) with 

 Hyale nigra Haswell (1880) on the basis of Schellenberg's (1928) 

 review of the latter. Further study of H. frequens in comparison with 

 H. rubra (Thomson), as reviewed by Hurley (1957), reveals such a 

 close identity that it is proposed to establish H. frequens as a sub- 

 species of that entity. Hyale nigra also may be identical to H. rubra. 

 Indeed, H. nigra is not well known, not having been reviewed from 

 its type area, east Australia, except presumably by Stebbing (1906) 

 who added sufficient detail to make one believe he saw specimens of 

 the species. Schellenberg (1928) also examined specimens of H. nigra 

 from Australia. That slight differences of male gnathopod 2 occurring 

 between H. rubra and H. nigra, as stated by Stebbing, are valid, is 

 tempered by the knowledge that mounting positions of the appendage 

 can vary. 



The distinctions between individuals of the Californian Hyale 

 presumed to be H. frequens Stout, and Hurley's figures are minor 

 but of sufficient degree to warrant subspecific allocation. The indi- 

 viduals are contrasted to a single male collected at Cedros Island, 

 Baja California, reported above, which is assigned to Hyale rubra 

 rubra. 



Hyale camptonyx (Heller), as figured by Chevreux (1911) and 

 Chevreux and Fage (1925), is a difficult species to assess in light of 

 variations occurring in pereopod 5 and the maxillipedal dactyl by 

 various authors. That species must be clarified in relation to Hurley's 

 analysis of H. rubra. 



