186 U.S. NATIONAL MUSEUM BULLETIN 25 8 



Summary of remarks on Lysianassa. — Interspecific intergrada- 

 tions of upper lip, epistome, uropod 3, uropod 2, gnathopod 2, maxil- 

 lipeds, and maxillae 1-2 prevent use of their alternate extreme 

 conditions as generic characters in the Lysianassa-complex. Inter- 

 gradation has been demonstrated also in the length of article 1 of 

 the mandibular palp and there is little qualitative value in the mandi- 

 bular molars. Some species of Lysianassa may have to be segregated 

 generically because of their second maxillae (L. hypocrita), but there 

 appears to be sufficient variation in the length of the fourth maxilli- 

 pedal palp article to render its generic significance useless. The 

 elongation of the second antennal flagellum in males of Aruga may 

 have subgeneric value as has been suggested for Harpinia and Har- 

 piniopsis in the Phoxocephalidae by J. L. Barnard (1958) and for 

 Aruga and Lysianopsis by J. L. Barnard (1955c). The writer suggests 

 a need for provisional fusion of all of the genera as listed in the syn- 

 onymy at the beginning of this discussion in anticipation that Lysia- 

 nassa costae, the type species, will fall into the scheme as developed. 



If variations in articulation of outer rami of uropod 3 are to be 

 ignored in the group of genera placed in the new subfamily Lysianas- 

 sinae by Hurley (1963), as well as the constriction of the inner ramus 

 of uropod 2, the generalized shape of maxilla 2 and the condition 

 of the maxillipedal dactyl, then the distinctions of other genera 

 allied to Lysianassa must be justified in ways other than so con- 

 veniently arranged by Hurley. Each of the genera Nannonyx, Para- 

 waldeckia, Paralibrotus and Parambasia have been so distinguished 

 in previous paragraphs. 



Key to Californian Species of Lysianassa 



1. Pereopods 1 and 2 with an enlarged distal spine on article 6 . . L. macromerus 

 Pereopods 1 and 2 lacking an enlarged distal spine on article 6 2 



2. Epistome projecting as far as upper lip 4 



Upper lip projecting much farther than epistome 3 



3. Third pleonal epimeron slightly convex posteriorly L. holmesi 



Third pleonal epimeron posteriorly produced into a quadrate plate . L. oculata 



4. Inner ramus of uropod 3 one third as broad and about five eighths as long as 



outer ramus, peduncle strongly expanded laterally, outer ramus uni- 



articulate L. pariter 



Inner ramus of uropod 3 nearly as broad and about four fifths as long as 

 outer ramus, peduncle unexpanded laterally, outer ramus Particulate 



L. dissimilis 



Lysianassa dissimilis (Stout), new combination 



Figure 47 a-f 



Nannonyx dissimilis Stout, 1913, pp. 638-639. 



Aruga dissimilis: Shoemaker, 1942b, p. 7, fig. 2. — J. L. Barnard, 1955c, pp. 100- 



103, pi. 29, figs, g, i.— Gurjanova, 1962, pp. 297-299, fig. 97.— J. L. Barnard, 



1964a, p. 230. 



