74 



UNITED STATES NATIONAL MUSEUM BULLETIN 22 4 



19. First posterior cell open 20 



First posterior cell closed and stalked ; mesonotal pile setate, 

 flat appressed Perasis Hermann 



20. Short, fine appressed setate flies of a bare, denuded appear- 



ance. Small, brown and grey species with short, robust, 

 apically rounded abdomen. Face with short, scattered 

 pile and the short proboscis is shorter than the face. 

 Female terminalia concealed by cupped terminal segment. 



TOKEBROMA Hull 



Flies of moderately long, fine pile; abdomen more elongate; 

 face below with a row of stout bristles, proboscis longer 

 than face Laphystiella, new subgenus 



21. PuMlli absent or reduced to minute stubs 22 



Pulvilli present 23 



22. Marginal cell closed with a long stalk. End vein of discal 



and fourth posterior cells nearly or quite aligned. Ex- 

 tremely bare, microsetate flies (Laphriinae). 



Antpodetds Hermann 

 Marginal cell closed with a short stalk, if any. End of 

 second vein strongly recurrent. End vein of discal and 

 fourth posterior cells not at all aligned. Pile dense and 

 short setate but not so contrastingly bare. Oral and oc- 

 cipital bristles spikelike. Last tarsal segment swollen 

 and spiculate below Macahtba Carrera 



23. Second vein not recurrent. Hind femur slender, its pile 



and bristles weak Laphtstia Loew 



Second vein moderately to strongly recurrent at apex. 

 Hind femur except in Martinia usually large, or very 

 large and thickened, and usually with strong, spine bear- 

 ing, ventral tubercles 23a 



23a. Second vein moderately recurrent at apex. Hind femur 

 slender, with at most weak pile and bristles. Pulvilli 

 nearly as long as claw but quite slender. Scutellum 



with bristles Martinia, new genus 



Second vein strongly recurrent. Hind femur large, usu- 

 ally with spine bearing tubercles below. Scutellum with- 

 out bristles ; margin and disc with short, stiff, usually 

 curled pile. Often rather large and robust flies . . 24 



24. Ambient vein complete. Hind femur quite stout, usually 



with strong tubercles and spinous bristles. Male termi- 

 nalia enlarged ventrally . . . Hoplistomerus Macquart 

 Ambient vein ends at the anal cell; smaller flies. Hind 

 femur enlarged but not exceptionally. Weak femoral 

 tubercles present, or absent altogether. Male terminalia 

 not unusually prominent ventrally . Trichardis Hermann 



Genus Laphystia Loew 



Figures 30, 443, 881, 890, 1978, 1980, 2005, 2007, 2022 



Laphystia Loew, Linnaea Entomologica, vol. 2, p. 538, 1847. 

 Type of genus: Laphystia sabulicola Loew, 1847, by 

 monotypy. 



Asicya Lynch Arribalzaga, Anal. Soc. Cient. Argentina, vol. 9, 

 p. 224, 1880. Type of genus : Asicya fasciata Lynch Arri- 

 balzaga, 1880, by monotypy. 



Laphyctis Loew, Ofvers. Svenska Vet.-Akad. Forhandl., vol. 

 15, p. 338, 1859. Type of genus: Stichopogon gigantella 

 Loew, 1852, by original designation. 



Laphystia has 1 subgenus, Laphystiella, new 

 subgenus. 



Pollinose, medium size to small flies, of robust type, 

 characterized by the generally bare appearance of 

 thorax and especially the abdomen, which is due to the 

 short, appressed pile. The robust third antennal seg- 

 ment bears a very short microsegment, followed by a 

 larger, oblique, spoon-shaped microsegment. The 

 cylindrical proboscis is unusually short and obtuse 

 apically. The face and slightly divergent front are 

 both short and wide. On the wing the anterior brancli 

 of the third vein is strongly sigmoid and arched at the 

 base; the first posterior cell is narrow and narrowly 

 open, or rarely closed in the margin and the marginal 

 cell varies from moderately widely open to closed in 

 the margin. Moreover, the sides of the tergites bear 

 lateral, subposterior rows of distinct bristles. Length 

 8 to 15 mm. 



Head, lateral aspect : The length is two-thirds of the 

 height. The face is moderately prominent throughout, 

 nearly straight and vertical. The eye is strongly con- 

 vex anteriorly and usually equally developed above and 

 below. In the type of genus there is a gradual reces- 

 sion of the eye beginning at the upper third and con- 

 tinuing below. The occiput is moderately prominent 



but obliterated near the vertex and is more conspicuous 

 ventrally. Pile of occiput abundant, rather long and 

 fine, especially below, but short and scanty dorsally. 

 The upper occiput has 3 to 5 pairs of bristles, some- 

 times reduced to thick hairs or in other species they 

 are stout and sharply differentiated. The proboscis is 

 shorter than the face in the type of genus, obtuse 

 apically, cylindrical on the outer half, and very 

 strongly and abruptly swollen at the base; in some 

 species it is longer than the face. A medial ridge is 

 present; the apex has fine bristly hair dorsally and 

 along the apex a middle lateral fringe which may be 

 turned downward. Base of proboscis with a number 

 of long, fine hairs. Palpus minute in the type of 

 genus ; first segment short and swollen, second segment 

 slender, short and cylindrical, bare, porate, with 1 or 

 2 fine apical bristles; the first segment is hemicylin- 

 drical, excavated and sometimes more or less fused. 



Head, anterior aspect : Face below antenna one-third 

 the head width and slightly divergent below. Face 

 wholly pubescent, with rather abundant long pile lat- 

 erally, becoming longer above the epistoma and with 1 

 or 2 pairs of distinctly slender bristles in the middle 

 above the epistoma, or there may be two rows of stout, 

 long bristles containing 6 or 7 pairs. The antenna is at- 

 tached at the upper third, elongate but rather stout, the 

 first two segments of nearly equal length, the second a 

 little shorter. The third segment is nearly twice as long 

 as the first two and of nearly uniform thickness 

 throughout; it is rather thick and bears apically two 

 microsegments ; the first is quite short, the second is 

 longer but extended outward obliquely ventrally so that 

 the apex is strongly truncate, almost spoonlike with 

 enclosed minute concealed spine. The pile of the first 

 antennal segment consists of stiff, bristly hair below, a 

 few setae above and characteristically with 2 pairs of 

 stout, long, anteriorly directed bristles ; second segment 



