ROBBER FLIES OF THE WORLD 



463 



ker (1862) ; grandis Macquart (1838) ; interponens 

 Walker (1861); noscibil'is Austen (1915); raptor 

 Austen (1915) ; rufpes Macquart (1848) ; tastnanensis 

 Macquart (1847) ; transacius Walker (1864) ; triuin- 

 phansBezzi (1928). 



Country unknown: Promachvs alhibarbis Macquart 

 (1838); castanipes Macquart (1838); dorsalis Mac- 

 quart (1838); geminus Walker (1849); incisus Mac- 

 quart (1838); longiterehratus Macquart (1838); 

 'imoerens Wiedemann (1828) ; porrectvs Walker 

 (1851); ntf-pes Macquart (1838); separatus Walker 

 (1855) ; tibialis Macquart (1838) ; vicinus Macquart 

 (1838). 



Malloch (1916, 1917) contrasts the pupae of Pro- 

 machiis fitchii and vertebratus. 



Engel (1930, 1932) was the first person to divide the 

 genus Provmchus into subgenera. In his 2 papers the 

 genera Philomachus Karsch and Trypanoides Becker 

 were treated as subgenera and I believe he was entirely 

 correct in his approach. Engel also created a new sub- 

 genus, Etiagaedium Engel, characterized by a peculiar 

 ovipositor. Parapromachus^ new subgenus for Proma- 

 chus leoninus is separated by the absence of bristles on 

 face and scutellum and the short, stout, robust ab- 

 domen and less shaip claws. 



The Promachus complex is separated from the Nerax 

 group of genera by the absence of any extensive gib- 

 bosity on the face as well as the character of vena- 

 tion ; the venation allies it to Mallophora Macquart to 

 which it is comiected by Promachina Bromley and 

 Paraj)ro7nachus, new subgenus. The blimt claws and 

 reduced male and female terminalia separate Mallo- 

 phora from Promachus. 



Hobby (1936) has suggested on what are slender 

 grounds the use of the name Bactria Megerle, which 

 was a manuscript name Megerle added by brief men- 

 tion only to the end of the description by Meigen ( 1820 ) 

 of Asilus picfus, and without any differentiation or dis- 

 tinguishing characters mentioned. Kertesz (1909) 

 though aware of Bactria rejected it and though Co- 

 quillett (1910) adopted it, the name has been rejected 

 by nearly all authors. On the principle of conserva- 

 tion of nomenclature, I have followed Engel in using 

 Promachus. Eicardo (1920) suggests other synonymy. 

 See addendum (p. 595). 



4. Male terminalia large and prominent and generally with a 



conspicuous dorsal tuft of white or silvery pile. 



Promachus Loew 



Male terminalia small, reduced in size and generally without 



conspicuous, white tufts of hair 5 



5. Second antennal segment twice as long as wide; margin of 



subepistoma with 4 or 5 quite stout bristles ; palpus slender. 

 Superior forceps elongate, arched outwardly and recurved 

 at apex, hollowed medially, leaving a conspicuous internal 

 cavity ; aedeagus tubular, extended beyond forceps. 



E.VAGAEDIUM Engel 

 Second antennal segment as long as wide ; margin of epistoma 

 generally with coarse, long hairs or slender, bristly hairs; 

 palpus stout. Superior forceps elongate, its inner dorsal 

 margin not arched, the cavity enclosing proctiger not 

 enlarged ; aedeagus very short Trypanoides Becker 



6. Ovipositor short, composed of segments 8 to 10 ; the abdomen 



composed of segments 1 to 7 7 



Ovipositor elongate, slender, the component parts each rather 

 long and attenuate, and comprising segments 6 to 10 ; ab- 

 domen composed of segments 1 to 5 . . Trypanoides Becker 



7. Ninth segment, except the base, and the proctiger (tenth 



segment) extremely flattened and compressed laterally ; the 

 tenth wedged into the ninth quite as in Eutolmus Loew. 



Enaoaeditjm Engel 



Segments of ovipositor conical or cylindroid, the last segment 



slightly compressed ; tenth segment not wedged into the 



ninth Promachus Loew 



Subgenus Aniblyonychus Hermann 



Aml)lyo)i!/c}iHS Hermann, Archiv Naturgesch., AM. a., vol. 87, 

 p. 118, 1921. Type of subgenus : Promachus wiedemanni 

 Schiner, 18G7, by original designation. 



Hermann (1921) listed a number of species of Asilids 

 and Mydaids from Paraguay and among these proposes, 

 without characterization, the name Amhlyonychus as 

 subgenus of Promachus Loew with Promachus wiede- 

 manni as type of genus. Since the name is obviously 

 derived from the Greek, amhlys., meaning blunt, obtuse 

 or dull, with reference to the claws, I suspect that 

 Hermann has here anticipated Bromley (1934) in the 

 erection of the genus Promachina Bromley, and we may 

 have to use Hermann's name. Since I have not seen 

 specimens of Schiner 's species I refrain from any defi- 

 nite determination as to the status of these names. It 

 should be remarked that Promachina is a rather distinct 

 New World genus. 



KEY TO SUBGENERA OF PROMACHUS 



1. Apex of antennal style conspicuously dilated. 



Philomachus Karsch 

 Apex of antennal style pointed or at most slightly flattened . 2 



2. Short, stout species ; the claws stout and only moderately 



sharpened. The ambient vein ends at the beginning of the 

 anal cell ; scutellum thick, long pilose, without bristles. 



Parapromachus, new subgenus 



Ambient vein complete; claws .slender, sharp; not short, 



robust species ; scutellum usually with bristles and pile . 3 



3. Males 4 



Females 6 



Subgenus Enagaediutn Engel 



Figures 313, 795, 1380, 1389, 2192, 2215, 2423, 2424 



Enagaedium Engel, Konowia, vol. 8, p. 459, 1929. Type of 

 subgenus : Asilus poetinus Walker, 1849, by original desig- 

 nation. 



Comparatively small, rather slender flies with a dis- 

 tinctive ovipositor, which is very similar to the ovipos- 

 itor of Dysmachus Loew and Eutolmus Loew, and of 

 which it may be considered a parallelism. The ninth 

 and tenth segments are quite flat appressed laterally 



