490 



UNITED STATES NATIONAL MUSEUM BULLETIN 224 



Quickly distinguished from Proctacmithus Macquart by 

 the widely dihited second submarginal cell and the 

 abruptly widened first posterior cell, together with the 

 evanescent posterior veins and ambient vein. Female 

 ovipositor with only dense, stubby setae. Length 35 to 

 46 mm. 



Head, anterior aspect : The head is of medium length, 

 the face is plane, quite short above and more conspicuous 

 below, only because of the anteroventral recession of the 

 eye, which is gradual and leaves the eye quite narrowed 

 below. The occiput is moderately developed through- 

 out and bears stout bristles extending to the bottom of 

 the eye and to the vertex above. The proboscis is rather 

 slender, elongate, somewhat attenuate on the discal 

 fifth and has fine, apical pile. Palpus of one segment, 

 comparatively short and small and with a trace of the 

 basal segment. The antenna is attached at the upper 

 third of the head and rather elongate ; the third segment 

 is as long as the combined length of the first two, bears 

 a distmct, short microsegment and a thick, spine-tipped 

 style, as long or longer than this segment. 



Head, anterior aspect: The head not very wide but 

 little wider than high. The face below the antenna is 

 less than a fifth the head width, slightly divergent over 

 the middle and more strongly below; the cheeks are 

 prominent. The face is densely pubescent, without pile 

 on the upper third and with a high medial triangle of 

 quite stout, long, white bristles on the lower half of the 

 face, mixed with short, slender bristles or bristly pile. 

 There are 3 pairs of long, stout bristles along the sides 

 of the subepistomal region. Front short, pollinose, 

 sunken, with an ocular and subocular row of short 

 bristles and bristly pile ; its sides are slightly divergent 

 and narrowed again at the vertex. Vertex moderately 

 excavated ; the ocellarimn is low, with 3 pairs of bristles 

 between the ocelli and 1 pair beliind. 



Thorax: The mesonotum is long, densely pale pol- 

 linose and densely covered with fine, subappressed setae. 

 Acrostical elements are undifferentiated and dorsocen- 

 tral elements present only at the anterior plane of the 

 postalar region, where there are 3 or 4 pairs of long, 

 stout, bristles. Humerus with short setae, the lateral 

 bristles are stout, mostly quite long and consist of 1 or 

 2 small posthumeral, 2 notopleural, 1 supraalar, 1 post- 

 supraalar, 1 to 2 suprapostalar, 3 or 4 postalar, and 3 

 pairs of scutellar bristles. The scutellum is densely 

 pollinose, and thick only basally, and it has a trace of 

 an impressed rim; the disc bears scattered, fine pile. 

 Pronotum with 15 to 20 pairs of stout, spinous bristles; 

 the lateral propleuron also with stout bristles. Ptero- 

 pleuron and posthypopleuron each with 2 or 3 conspicu- 

 ously stout bristles. JNIetanotal callosity with a dense 

 patch of conspicuous, coarse pile; prosternum fused. 

 Metastemum pilose laterally and below, and the post- 

 metacoxal area membranous. Posterior basalare with 

 bristles and stiif ha ire ; anterior basalare pollinose only ; 

 tegula with veiy fine hairs. 



Legs: The legs are stout, the femora elongate, but 

 scarcely or not at all swollen. The pile of the legs is 

 dense, coarse, and plastered flatly upon the surface. 



especially on the hind pair. Bristles are quite stout, 

 only moderately long, and not abimdant. All the coxae 

 laterally bear stout bristles. The hind femur bears 2 

 or 3 lateral bristles, a transverse, dorsal pair at the 

 outer sixth, 2 ventrolateral, 2 lateral, and 5 medial 

 bristles all at the apex, and in addition there are 5 ven- 

 trolateral and 3 or 4 ventromedial bristles. The hind 

 tibia bears 2 dorsolateral near the middle and 2 dorsally 

 nearly opposite, besides 3 short, basal, and 4 long, distal 

 ventrolateral bristles. Middle femur with 2 stout, an- 

 terior bristles near the middle, 2 dorsomedially at the 

 apex, a ventromedial row of 5, and a ventrolateral row 

 of 2 bristles. The middle tibia usually has 2 or some- 

 times 3 stout, anterior, apical bristles and 4 or 5 spike- 

 like, ventral bristles on the basal half. Anterior tibial 

 bristles are similar, with both dorsal rows rather stout 

 and long, containing 3 elements and rarely 4 elements, 

 and with 2 posteroventral bristles. Claws stout, mod- 

 erately sharp ; the pulvilli and the bladelike empodium 

 long. 



Wings: The wings are comparatively short and 

 broad, much shorter than the abdomen ; the second mar- 

 ginal cell is widely dilated over most of its length, but 

 arises gradually. The second posterior cell, however, is 

 greatly widened and arises abruptly. At least the pos- 

 terior veins are completely evanescent and the whole of 

 the ambient vein likewise. Fourth posterior cell closed 

 with a long stalk; the alula large. There is a strong 

 similarity between the venation and that of Nearatus 

 Ricardo and also of Blepharotes Westwood. 



Abdomen: The abdomen is elongate, conspicuously 

 tapered, especially in the female, but more gently in 

 males, because of the large clublike terminalia. Pile is 

 moderately abundant but flat appressed and setate and 

 scarcely longer laterally, even on the basal segment. 

 Sides of the first tergite with about 10 long and many 

 shorter, stout bristles ; middle of second tergite with a 

 patch of 6 stout bristles and some smaller ones and its 

 hind margin has 7 or 8 pairs. Third tergite with 

 lateral and also postmarginal bristles and the fourth 

 tergite only with postmarginal bristles. Second to 

 fourth sternites with stout bristles. Males with eight 

 tergites; the eighth is half as long as the seventh. Fe- 

 males with seven segments not incorporated in the ovi- 

 positor and the seventh is partially modified. The male 

 terminalia are short but large and clublike. The su- 

 perior forceps are especially stout, the gonopod much 

 shorter, the hypandrium well developed, the proctiger 

 large and oblique. Female ovipositor elongate and 

 cylindrical and somewhat conical. Most of the length 

 is contributed by the eighth segment, which is nearly 

 thre« times the combined length of the last two seg- 

 ments. The tenth segment both above and below bears 

 dense, short, stubby setae. 



Distribution: Palaearctic: Satanas agha Engel 

 (1934) ; chan Engel (1934) ; fuscanipennis Macquart 

 (1855) ; gigas Eversmaim (1855) ; minoi' Portschinsky 

 (1887) ; niveus Macquart (1838) ; shah Rondani 

 (1873) ; testaceicomis Macquart (1855). 



