82 U.S. NATIONAL MUSEUM BULLETIN 29 



as much larger and reaching the middle of the sides of the elytra. 

 Schaeffer (MS. key) described the humeral spot in quadrujplex as 

 small and scarcely wider apically and in that of quadriguttulus as 

 larger and obliquely widening toward the apex. In the USNM collec- 

 tion, there are over 400 specimens previously determined as qiiadru- 

 plex or quadriguttulus. These series show considerable variation in 

 the development of the basal elytral spot. Within the eastern and 

 middle parts of the range of the specimens, I find continuous inter- 

 mediates between individuals representing the extremes in develop- 

 ment of the basal spot. I have synonymized quadriguttulus because 

 these color differences are not consistent and intermediate forms are 

 numerous. At one extreme of development, the spot is within the basal 

 one-third of the elytra, and its inner margin borders the fifth interval. 

 At the other extreme, the spot is within the basal half of the elytra, 

 its inner margin is diagonal at its base, joins the fourth stria, and 

 apically attains the middle of the second interval. Most of the in- 

 dividuals with the basal spot reduced are from the New England 

 States (New York, Massachusetts, Rhode Island) but some extend 

 to Michigan, Wisconsin, and Illinois. None of these individuals bear 

 labels indicating they were collected in the westernmost part of the 

 range (Iowa, Nebraska, Kansas). The specimens from the western 

 part of the range have the spot at about its maximum size. In these 

 individuals, the inner margin of the basal spot is nearly always 

 straight. Individuals from other parts of the range that have a large 

 spot often have the inner margin arcuate, and the markings thus 

 are similar to those of b. binominis. 



Douglass (1929, p. 8) recorded this species from Rhus glabra L. 



Distribution. — The USNM collection has a total of 403 individ- 

 uals from Ontario, New Hampshire, Connecticut, Rhode Island, 

 Massachusetts, New York, New Jersey, Delaware, Maryland, Penn- 

 sylvania, Virginia, North Carolina, Alabama, Georgia, Kentucky, 

 West Virginia, Ohio, Indiana, Illinois, Michigan, Wisconsin, North 

 Dakota, Arkansas, Missouri, Louisiana, Iowa, Kansas, Nebraska, 

 Montana, Minnesota, Oklahoma, and Texas. 



Cryptocephalus quercus Schaeffer 



Figure 45 

 Cryptocephalus quercus Schaeffer, 1906, p. 232. 



Pronotum: DuU light orange to orange, disk usually vaguely 

 darker; punctation usually dual, larger punctures minute to blending 

 with small punctures, latter sparse. 



Elytra: Dull creamy yellow to light orange, mth striae, humerus, 

 and dark markings more or less reddish. Each elytron with two vague 



