GAMMARIDEAN AMPHIPODA 81 



native nomenclatural solution would be to retain all tmetonyxes in 

 Tmetonyx and base the genus on the presence of a dactylar tooth 

 regardless of the condition of the first coxa. This would require 

 the transfer of several species now assigned to '"^TryiylioscC into 

 Tmetonyx because of their dactylar teeth. The problem is compli- 

 cated by the occurrence of several species with intergrading conditions 

 in the size of dactylar teeth and this suggests that the nomenclatural 

 valuation lies at the subgeneric level. The whole tryphosa problem 

 is also complicated by the uncertainty of classification in related genera 

 as discussed in the introduction to the Lysianassidae, under Aiiibas- 

 iopsis fo7nes and Uristes perspinis and in the discussion of the relation- 

 ships of the new "tryphosa" below. 



Tryphosella Bonnier {=''^Tryphosa'^^ auct.) 



Tryphosella Bonnier, 1893, pp. 170-171. 



"Tryphosa".— Sam, 1S95, pp. 75-76.— Stubbing, 1906, p. 68.— J. L. Barnard, 

 1962b, pp. 28-29 (not Boeck, 1871; but including all species except the 

 type-species of Tryphosa, Anonyx nanus Kr0yer). 



"Tmetonyx". — auct. (not Stebbing, 1906, but including all species except type- 

 species, Oniscus cicada O. Fabi-icius). 



ILepidepecreopsis Stepbensen, 1925, p. 119. 



Type-species. — Trypliosella sarsi Bonnier (present selection). 

 This species is identified as '■''Tryphosa nana''' of Sars, 1895, p. 76, pi. 

 27, fig. 1 (not Kr0yer) . 



Diagnosis. — Lysianassid with "large" head, lateral cephalic lobes, 

 normal-sized and noncoalesced thoracic and abdominal segments, fully 

 developed mouthparts and uropods, accessory flagellum multiarticu- 

 late; mouthparts grouped in quadrate bundle below head; prebuccal 

 parts strongly developed, epistome protuberant and dominating small 

 lobe of upper lip, occasionally epistome subacutely produced; mandib- 

 ular incisor medially smooth, molar of medium size or small, not 

 cuboidal, asymmetrical, occasionally slightly attenuated but never 

 conico-laminate, strongly setulose; maxilla 1 with 2-articulate palp, 

 outer plate with medium-sized (not elephantine) spines, inner plate 

 with a few unmodified setae; maxilla 2, lower lip and maxilliped of 

 ordinary dimensions; coxa 1 slightly shoi-tened, tapering distally and 

 slightly concealed by coxa 2 ; coxa 2 large and quadrate ; giiathopod 1 

 subchelate, no articles grossly enlarged ; dactyl often bearing accessory 

 marginal tooth; inner ramus of uropod 2 simple; outer ramus of 

 uropod 3 biarticulate, inner ramus of normal elongation ; telson cleft 

 distinctly. 



Remarks. — Three subgenera might be established within this genus, 

 the subgenus Lepidepecreopsls being composed currently iof one 

 species with asetulose mandibular molar, the other 2 subgenera with 



