392 HYDRANGEACEAE 



Family 58. HYDRANGEACEAE. Hydrangea Family. 



Shrubs or trees, or rarelj' vines. Leaves opposite, without stipules. Flow- 

 ers commonly perfect, in cymes. Hypanthium well developed, usually 

 ribbed. Sepals 4-10. Petals as many. Stamens from 8 to many. Gynoe- 

 cium of 2-5, rarelj^ 10, united carpels. Ovary partly or wholly inferior; 

 styles distinct or united, sometimes wanting. Fruit a capsule. 



Caosule obovoid or obcouic, the beaks lising abruptly from the body. 



1. Philadelphxts, 



Capsule conic or ovoid, the beaks tapering gradually from the body. 

 Filaments and anthers unappendaged ; sepals and petals 5. 



Leaf-blades toothed; hypanthium short during anthesis. 2. Edwtnta. 



Leaf-blades entire; hypantluum long during anthesis. 3. FENDlEREliLA. 



Filaments appendaged under the anthers; anthers appendaged at the top: sepals 



and petals 4, 4. Fendlera. 



1. PHILADf LPHXJS L. Mock Orange, SYRmoA. 



Branching shrubs. Leaves toothed or entire. Flowers perfect, borne at 

 the ends of short leafy branches, solitary or in cymes. Sepals 4, rarely 5, valvate, 

 persistent, more or less tomentose within. Petals 4, rarely 5, convolute, white 

 or ochroleucous. _ Stamens usually many, 25-60; filaments flat, subulate, free 

 or more or less united below; anthers short, didymous. Ovary at least two-thirds 

 inferior and adnate to the hypanthium, 4-celled; styles elongate, distinct or more 

 or less united; stigmas distinct or more or less united. Ovules numerous, pendu- 

 lous. Capsule more or less woody, loculicidal, the septa separating in the cen- 

 ter. Seeds numerous; coat reticulate; albimien fleshy. 



Leaf-blades thin, not leatherj". usually toothed or denticulate, raore than 3 cm. long; 



flowers several. 



Petals ovate-lanceolate, acute or obtuse at the apex. 1. P. Helleri 



Petals oblong to rounded-o val ; rounded at the apex. 



Leaf-blades 3-5-ribbed, the lateral ribs or if 5-ri))bed the second or stronger pair 



meeting the midrib some distance from the base of the blade. 

 Bark of the second season red or chestnut-brown, with conspicuous cross- 



cracks, at last peeling off. 2. P. LervisiL 



Bark of the second season gray or yeUowish, neither cross-cracked nor peeling 

 _ , ^,^T' 3. P. columbianus. 



Leaf-blades 5-nbbed. all ribs originating at the base of the blade. 

 -,,,-„ 4. P. Gordonianus, 



Lear-blades smau, 0.5-2 cm. long, entire; flowers mostly solitary. 



Hypanthium 4-5 mm. high. 5. p. microphyllus. 



H>-pant:hium about 2 mm. high, or in fruit 3-4 mm. high. 



Styles wholly united or nearly so; stigmas sometimes also partly united. 



-^ - ^. . ^ , 6- P. occidentalis . 



Styles distmct for at least half their length; stigmas distinct. 



7. P. minutus. 



1- P. Helleri Rydb. Shrub 2-3 m. high, with ascending branches; bark of 

 the young branches brov^-nish, glabrous, and of the second year yellowish or gray, 

 not exfoliating; leaf-blades ovate or hince-ovate, denticulate or rarely entire, 

 4r-7 cm. long, glabrous, or sparingly hairy along the veins beneath; flowers 5-11, 

 falsely racemose; sepals 7-8 mm. long, short-acuminate; petals white, 15-18 mm. 

 long; styles united to near the apex. Hills: Ida. SuhmonL Jl. 



2. P. Lewisii Pm-sh. Shrub 1-2 m. high, with ascending branches; bark 

 of the yoimg t^igs red, glabrous; leaf-blades ovate-lanceolate or ovate, glabrous 

 above, hairy on the veins and in their axils beneath, entire or denticulate; sepals 

 lance-ovate, abruptly acuminate; petals white, 10-15 mm. long; styles united 

 half their length; capsule about 1 cm. long. (?) P. intermedius A. Nels. River 

 ^—'- and hills: B.C.— Mont.— Ida.— Ore. SubmonL~M onL :My-Jl. 



3. P. columbianus Koehne. Shrub 2-3 m. high, vnth ascending branches; 



bark of the young tvngs mostly yellowish, glabrous or pubescent; leaf -blades 



lanceolate to ovate, more or less acuminate, denticulate or entire, more or less 



hair>' on both sides; flowers 3-9, falsely racemose; sepals lance-ovate, acuminate; 



petals white, 15-20 mm. long; styles united half their length; capsule about 12 



mm long, somewhat 4-angled below. River banks and copses; B.C.— Ida.— 

 Calif. SuhmonL My-Jl. 



