324 KOFOID AND SWEZY. 



have separated (PL 3, Figs. 26-30) and seems to act as an elastic 

 tether which turns the blepharoplast poles of the daughter nuclei 

 towards one another from their original separation of 180° (PL 3, 

 Figs. 26-29) when the protean activity of the organism results in a 

 wider separation of the nuclei. In Figure 28 the paradesmose has 

 almost its original length and position and the nuclei are but slightly 

 turned. In Figure 29 the nuclei have parted for a distance equal to a 

 nuclear diameter, the paradesmose is very thin and straight as though 

 under tension and both nuclei are turned as though drawn by their 

 blepharoplasts. In Figure 26 the nuclei are nearly two nuclear 

 diameters apart but the blepharoplast of the nucleus to the right has 

 become detached from its nucleus and the paradesmose has shortened 

 up and thickened as though it had contracted. 



It can no longer be detected between the blepharoplasts after the 

 axostyle has begun to divide. It fades away without leaving any 

 special organelle as its successor. It does not form the daughter 

 axostyles as described by Prowazek (1904) and by Dobell (1909). 



The Anaphase. 



The anaphase in which the chromosomes migrate to the poles of 

 the spindle and the nuclear membrane constricts is of some duration 

 for a number of cells in this period have been found in our preparations. 

 As the chromosomes approach the poles of the spindle the spindle 

 fibres grow thicker and darker or the chromatin material of the chro- 

 mosomes flows out towards the blepharoplasts until they seem to be 

 actually in contact with it (PL 3, Figs. 24-28) for a time. During 

 this period the chromosomes retain their individuality to a striking 

 degree. 



The daughter nuclei are formed by the constriction of the nuclear 

 membrane in the equator of the broadly fusiform spindle (PL 3, Figs. 

 24, 25). Nuclei in this stage of constriction are very rarely seen in 

 preparations. This process is therefore quickly accomplished. 



The Telophase. 



The telophase, in which the nuclei return to the vegetative or 

 "resting" condition, is accomplished by the detachment of the chro- 

 mosomes from the blepharoplast pole of the nuclear membrane (PL 3, 



