TRICHOMONAD FLAGELLATES. 355 



of all of the organelles. The following comments are made on his 

 conclusions with regard to the lower flagellates related to Tricho- 

 monas. 



The transfer of Devescovina to, and the inclusion of Foaina with the 

 Polymastigina is in accord with our conclusion that the chromatic 

 basal line of Trichomonas is the homologue of the parabasal in other 

 forms. His inference that the parabasal of Devcscov-ina divides at 

 mitosis is not supported by evidence of actual division. It grows out 

 independently of the old parabasal in Trichomonas and may be ex- 

 pected to do so in Devescovina. He still revives the old figure of 

 Trichomonas showing the stout body which he calls parabasal though 

 recognizing its rarity and temporary status, but does not reinvestigate 

 material nor recognize the relationships of this organ to the chromatic 

 basal rod or true parabasal. His evidence presented for the origin 

 of the new parabasal by outgrowth in the case of Stcphanonympha 

 is on the other hand amply conclusive. 



The extrusion of bacteria-like granules from the nucleus at the 

 telophase in Dcrescotina is apparently analogous to the formation in 

 Trichomonas of extranuclear chromidia at the prophase as we haxe 

 described it. 



His description of the behavior of the extranuclear organelles in 

 mitosis in Devescovina differs in certain very important particulars 

 from our conclusions in the case of Trichomonas. In the first place 

 he figures (his pi. 13, figs. 10, 11) two extranuclear division centers. 

 One of these spins out the extranuclear paradesmose (his "Spindel"). 

 between the daughter division centers each of which consists of a 

 centriole to w'hich is attached a single parabasal and a group of flagella 

 exactly as in Trichomonas. These daughter centers become the ble- 

 pharoplasts of the daughter organisms. He states however that these 

 centers do not arise from the blepharoplast but independently of it, 

 no source being stated or evident in the figures. We believe this 

 inference incorrect and that they will be found to arise from the 

 parent blepharoplast. A second center, figured but twice, consists 

 of a dark granule within a light halo, both of which structures are spun 

 out in line parallel to the paradesmose. Their fate is unknown, but 

 he suggests that they become the suspensory lamellae coming from 

 blepharoplast past the nucleus towards the axostyle. This second 

 division center is regarded by him as the true blepharoplast and the 

 line between them as the " Desmose." Our interpretation of this 

 organelle, based on Trichoiuonas, is that this second dividing center 

 is not the blepharoplast at all but may perhaps be the remnant of the 



