MENDELISM AND OTIU-K MKn.OT>S OF DESCENT 195 



the ..amete parents, but must wait for them to btuld up a new 

 otr.isn,, Jhiclr is not a true cross or product of co„,ugat,on, 

 bu' a rep.;ductive by-product, so to speak, which has been .„ter 

 calated into the experiment which we or.g.nally intended to per- 

 form The further progress of our desired conjugation is mean- 

 while suspended, and remains in abeyance until pi-eparations 

 are made for the production of the next generation of sex-cells. 

 The results we were seeking, as a means of testing the inher- 

 itance relations of the characters of the original parental organ- 

 isms, have not yet been attained. 



CONJUGATE ORGANISMS PRODUCE PERJUGATE GAMETES. 



The organisms built up by the parent gametes while in the 

 coniugate condition may be called conjugate organisms, or con- 

 jjates. The next generation may be termed the fcrjugatc geu- 

 eration to indicate that it has passed through conjugation. The 

 gametes which produce this generation are ferjugate gametes, 

 and are the first result of the completed conjugation of the ongi- 

 nal gamete parents. The organisms which these gametes build 

 UP ffford our first visible evidence of the relations which the 

 parental characters have assumed as a result of conjugation. 



What we have been calling the first generation of offspring 

 of such a cross is thus found not to be a true result of the cross, 

 but a structure built of double or conjugate cells which repre- 

 sent vegetative subdivisions of the gamete parents before heir 

 conjugation has been completed. What we have been cal Img 

 the second generation of the cross is really the first generation 

 of the organisms which represent the results of a true union or 

 completed conjugation of the original gamete parents. 



With reference to the next preceding generation each sexually 

 reproduced higher plant or animal is a conjugate, that is, it is 

 built up by conjugating gametes derive* from the preceding 

 generation. With reference to the second preceding generation 

 each individual is to be reckoned as a perjugate, that is, it rep- 

 resents the result of the completed conjugation of the gametes 

 derived from the second preceding generation. The relation 

 of an organism to the grandparent generation is distinct and 

 different from its relation to the parent generation. It is pro- 



