82 THE JOURNAL OF BOTANY 



geiious, globose-lens-sliaped, blackish-brown, opening witli a pore, 

 100 /A diam., at first covered, then erumpent ; texture thin, brownish. 

 Spores oblong-fusoid, when young eseptate and biguttulate, 

 7-9 X 2 ft, then 1-septate, with 2, 3, or 4 guttules, 12-lG x 2^-3 /x, 

 hardly at all constricted, faintly curved at times, hyaline, occasionally 

 with a mucous appendage at the end ; all these forms occurring in 

 the same pycnidium. 



On living leaves of Oxalis Acetosella. Dalrv, A^'rshire (Bo}^!). 

 July, 1919. 



Evidently an earlier state of the variety recorded in Journ. Bot. 

 1918, p. 318, as var. vermiformis. The latter was found by Mr. Boyd 

 in August 1918, about -4^ miles from Dairy. The link between the 

 two forms is furnished by the type specimens of Saccardo, which had 

 spores 17-20 x 3-3^ fi^ with three septa. By referring back to 

 no. 31-4, VliyUosticta Ojcalidis, and remembering that the spots on 

 which these various fungi occur are all of exactly the same character, 

 it will be seen that there is a strong suggestion of a series resembling 

 that of Sej)foria Chenopodii (see Journ. Bot. 1917, p. 34G). This 

 gradual change of spore-form and spore-size, as development advances, 

 has been overlooked by many observers in the past, and its recogni- 

 tion must react greatl}^ upon the conception of a " species " among 

 the Coelomycetes. 



Many of the younger forms showed no trace of colour in the 

 spores, whereas those of Saccardo and of the var. vermiformis both 

 had a faint j^ellowish tinge. The four guttules possessed by the 

 largest of the spores of the present gathering are a step towards the 

 formation of the lateral septa of the later forms : even in the variety 

 vermiformis there were a few spores which were 4-guttulate, but 

 only 1-septate. 



The occurrence of more than one form of spore upon the same 

 host and even upon the same " spot " is, of course, no proof of 

 genetic connection, although the " spots " may in many cases be 

 regarded as pure cultures : but, when the various forms occur in the 

 same pycnidium, it would require strong evidence to rebut the sugges- 

 tion of a common origin. 



AMPHORULA, gen. nov. 



Pyenidia immersa, solidiuscula, carbonacea. Sporulse ampulli- 

 formes, longirostratte, septatse, hyalinse. 



Genus KeUermanicB Ell. & Ev. (in Journ. Mycol, 1885, p. 153; 

 Sacc. Syll. x. 337) affine, sed forma sporularum pycnidioque solidiore 

 bene distinctum. 



346. Amphorula sachalinensis, sp. unica. 



Pycnicliis sparsis, depresso-globosis vel placentiformibus, 300- 

 750 ft diam., vix papillatis^ immersis, epidermide tectis eique prirao 

 arete adhairentibus, eandem postea poro orbiculari minutissimo delude 

 ainpliore v. rimiformi penetrantibus, postremo epidermide emortiia 

 desiliente superticialibus aut subinde cum eadem dilapsis, diu astomis, 

 atris, periodio crasso opaco sed stratum versus proliferum jmllidiore 

 circummunitis. Sporulis elliptico-fusoideis, superne in longum ros- 



