44 CAtJI'OUNIA ACAKK.sn "I SCIKNCKS. 



eoiiiiccting the various rirj^ans with tlic Ixuly-WMll ami ictniirc licic no particular nicii- 

 tioii. 



Ill till- olhcr flitcllar r^oniitcs we- liiul l>iiiiil> ol traii.sver.sc inusclcs wliii-h 

 stretch (liafionaily (fig. 118) across the body wall from the ventral part, almost ini- 

 iiu'<liat('ly hehnv the ventral ganglion, lo liic vicinity ol' liic onter seta-. The crxls of 

 these muscles penetrate between the lobes of the longitudinal muscular layer (fig. 

 lis, /. //(.) These muscles are not contiiiMons through the somite l)nt are grouped in 

 bands. 



Ventral pnpif/n (figs. 120, 125 to 131). Aigilophilus is readily distingui.shed 

 from other Califoi'nia EndrilidiC so far known, by its ventral papilla', occupying the 

 ventral side of the intersegmental grooves from the spermatheea U) the segments next 

 posterior to the clilellum. In Ai-fjilophi/ns ii/inijioratin* orudtnn we meet sometimes with 

 as many a.s 7 or 8 pairs, while in A. ni. jjn/iil/l/'ri we find as high a nuniln r in a single 

 median row, under the ventral ganglion. In no instance did T (ind a papilla on the 

 segment itself, all invariably occnricd rising from the groove between the .somites, 

 beinii' in other words iiitersci'-mental. From cross-sections it will be seen that the 

 papilla consists of two distinct jiarls, one extiTioi' ami latci-al, consisting of elongated 

 hypodermic su[)porting cells, which more or less fully enclose the interior main Ixtdy 

 of the papilla. Tliis central part consists of larger or smaller gland-like bodies, vary- 

 ing in size and number according to the size and age of the pa])iila, etc. In some 

 papilhe these bodies fill the larger ])art of the papilla, in others they are confined to 

 the bolt(Mn or very center of the organ. As to the nature of these organs 1 am, as yet, 

 somewhat in doubt, though they certainly must be considered as sen.sory organs 

 of some kind. Staini'd with osmic acid the bodies present in sections a darker center, 

 which appears of ganglionic nature, around which are grooped larger .sacs which 

 again are compo.sed of smaller, light-refractive granules, giving the idea of a reticu- 

 lated protoplasm. Numerous nerve fibres connect with those bodies, and evidently 

 penetrate to the gaglionic center. Long tube-like cells butt directly on these gland- 

 idous bodies, while others grooi)eil in lyundles penetrate between them connecting with 

 nerve ganglia. The transition between the supporting cells and the drainpipe cells 

 of the papilla is sudden and not gradual. In fact llic line of demarcation iietween the 

 two is (juite prominent. The suj)|)orting cells in the young papilla, entirely enclose 

 the papilla, while in the larger and full grown papilla, tlu'V are pushed towards the 

 side leaving the whole center to the drainpipe cells and the glands. 



When stained with safranine the nuclei of tlie glandular cells l)ecome prominent, 

 less so when staineil with luematoxylon. The nuclei vary greatly in size and ninuber. 

 In some of the glandular bodies there are from one to three nuclei, in otlu'rs none. In 

 sections there are always less nuclei than divisions (cells?) apparently lying on the toji of 

 the reticulated mass. The nuclei may be seen directly above the nerve center, but 

 generally lliey aic found onlsidc of it. ll seems as if these glandulous bodies are 

 modified agglomeralioii> of hypodeiinic glands. 



'i"he airaniiemenl ol ihe iilamlnlous bodies and the lube-like cells i> verv 



