148 JOURNAL OF ENTOMOLOGY AND ZOOLOGY 



The duct from the spennatotheca is small and ciliated. The 

 s])ermatotheca itself is a large, spherical sac, lined with long 

 (lark-staining, eolinnuar cells. These have large, oval nuclei 

 .just below the free surface and in the resting condition (in the 

 one specimen where the female organs were dormant) ap]iear 

 covered with a brown cuticle. In all the other specimens the 

 cells lining the spermatotheca were much elongated and pouring 

 forth a secretion. The spermatotheca was much more expanded 

 in the active than in the dormant specimen and the epithelium 

 over part of the surface usually appeared to be more or less 

 broken down, perhaps due to the excessive secretion. This 

 would make it appear that the spermatotheca is more than a 

 mere resting place for the spermatozoa, the function which is 

 ascribed to it by Alder and Hancock. The duct from the 

 external opening to the spermatotheca leaves very near the 

 opening of the duct from the spennatotheca to the oviduct. 

 This is at first small but becomes larger and more strongly 

 ciliated, and ends close to the opening of the penis in a small 

 opening surrounded by a heavy ring of muscle fibres. 



The oviduct in the dormant specimen appears as a tortuous 

 channel lined with short cilia, surrounded by a mass of connec- 

 tive tissue, in which lie numerous branching glands. These are 

 rather large sacs lined with dark-staining columnar epithe- 

 lium, the nuclei of which lie at the base of the cells. In the 

 dormant state these glands do not differ much from each other, 

 but in the active state they become very much changed in 

 appearance and differentiate into two types. One becomes very 

 much larger, the cells becoming full of a homogeneous secre- 

 tion and swollen to a length of 120 microns or more, the cell 

 outlines become very indistinct, and the whole mass stains very 

 faintly, although the nuclei stain very deeply. The other gland 

 is small and lies in the center of the mass, and farther from the 

 external opening than the other. It consists of branching tubes 

 lined with columnar cells about 80 microns long and with rather 

 a large lumen. These cells are sharply differentiated from 

 those of the other gland l)y the fact that they stain deeply 

 and the secretion is granular and is poured out into the lumen 



