302- PROF. G. ELLIOT SMITH ON THE 



bliglitost degree from tliat common mammaliau form, such as the braiu of Taplrm 

 exhibits, to the characteristic Prosimian or Primate condition. The jiersistence of the 

 narrow g-vrus between the intercalary and pai'acalcarine sulci, when the mechanical 

 conditions clearly ])i-edispose to the fusion custoraaiy in non-Primates, is yet another 

 testimony to the view tliat Chiromys is a retrograde Primate. 



The calcarine sulcus begins at a point S mm. lielow the splenium of the corjius 

 callosum and 2 mm. behind the hippocampal ti.ssare ; it jjroceeds obliquely upward and 

 backward to a ])oint 5 mm. behind and slightly below the level of the splenium : there 

 it bifurcates into retrocalcarine and paracalcarine branches. The former proceeds hori- 

 zontally backward and the latter vertically upward. In one instance the paracalcarine 

 is separated from the rest of the great cingular sulcus by a narrow submerged strip 

 (tig. 38) : in another case (fig. .38 a) there is also merely a very narrow strip separating 

 the sulci, l)nt the upper end of tiie paracalcarine is prolonged beyond the region of 

 approximation. This is the condition wdiicli Milne-Edwards (and Chudzinski after him) 

 represents as l)eing the usual form in the Indrisinse. 



The Braix of the Extinct Lemuroib Globilemur. 



In 1897 Dr. Eorsyth Major described * a })laster mould of the cranial cavity of the 

 large sub-fossil Lemuroid to which he had previously given the name Glohilcinur 

 Flaconfti. 



I Avould not have deemed it necessary to attempt to add anythiiig to the clear and 

 sufficient (/. e. considering the state of the material) account given by this excellent 

 anatomist, if his memoir liad not been recently criticised by lludolf Burckhardt. "Whilst 

 I Avas actually engaged in studying the plaster mould and the cranium from Avhich it 

 Mas made in the British Museum, my attention was called to the brief report of Burck- 

 hardt's criticism, Avhich was then just published f . I therefore carefully re-examined in 

 the light of these criticisms the crania and the casts of Glohllemur [and also of 2Icga- 

 ladapis, to which I shall refer later in this memoir]. Professor Burckhardt has just been 

 ood enough to send me a co2:»y of the full memoir %, of Avhich the above-quoted note 

 may be regarded as an abstract. 



The brain of Glohilcmur was considerably larger than that of any living Lemur. It 

 had large flattened olfactory bulbs projecting very slightly beyond the apex of the 

 hemisphere, as in the genus Lemur. In fact the whole of the exposed part of the 

 rhinencephalon closely resembles that of Lemur. 



1'he cerebral hemispheres (fig. 39) cover the cerebellum to a slightly greater extent 

 than in most members of tlie genus Ljemnr ; the obliquity of the postero-inferior margin 

 of the hemisphere is about the same as in Lemur (fig. 40). In respect to both of these 

 points, Burekhardt's drawings are misleading [op. cU. tig. 1, p. 231). 



So far as the shape of the hemispheres is concerned, this cast differs from that of any 

 of the larger Lemurs, such as Chiromys, Liidris, Froplthecus, and Lemur. It is least 



» Proc. Royal Soc. 1S97, pp. 40 & 47, pi. 5. tigs. 1, 2, and ;i. 



t 'TagoblaU dcs V. Litei'iuitionalfii Zdologcn-Cougresses,' ]!urlin, August liXIl, Nimimi'i- 4, 14lli August, 

 j ''Das Uehirn zweier subfussilcr Ricseulcmureii aus iladagasear,"' Aiiatomischer An/.eiger, Rd. xx. 1901, 

 pp. 2:29-232, fig. 1 , a & «. 



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