FROGS OF SOUTHEASTERN BRAZIL — COCHRAN 341 



bars. Underside dirty white, dusted with fine gray dots, darkest on the throat, 

 breast and femur. 



Head and body length: 14 mm. 



Remarks. — In his introduction, Alil states that his new genus in 

 many respects suggests Eleutherodactylus , from which it is set apart 

 by the entire lack of teeth. In the entirely toothless genera of the 

 family, it stands next to Batrachophrynus Peters, being distinguished 

 from it by the web of the hind foot. This web is present in Batracho- 

 phrynus, absent in Phrynanodus. 



When I examined the type and unique example of Phrynanodus 

 nanus in 1938, I noted its lilceness to Eleutherodactylus parvus in 

 having a similar triangular black anal patch (not mentioned by Ahl) 

 and pointed toes. The back of nanus was much more tubercular 

 than in parvus, however, and definite dorsolateral folds are lacking 

 in the latter. More material is needed from the type locality to see 

 whether the absence of vomerine teeth is indeed a permanent condition 

 in nanus. 



Specimen examined 

 BRAZIL: 



Rio de Janeiro: Itatiaia, south slope at 1,000 to 2,000 meters, NHMW 

 (type of P. nanus), Zerney, 1927. 



Genu8 Physalaemus Fitzinger 



1824. Bufo Spix (part), p. 47. 



1826. Physalaemus (not Physalaemus Fitzinger, 1843) Fitzinger, pp. 39, 65. 



(Genotype Physalaemus cuvieri Fitzinger = Liiiperus albonotatus 



Steindachner.) 

 1830. Paludicola Wagler, 1830a, p. 206. 

 1862. Rhinoderma Reinhardt and LtJxKEN, p. 171. 

 1862. Gomphobaies Reinhardt and LOtken, pp. 171-2. 

 1864. Eupemphix Steindachner, 1864a, p. 271 (part). 

 1864. Nattereria Steindachner, 1864a, p. 279. 

 1868. Lystris Cope (part), p. 312. 

 1875. Pleurodema Espada, p. 87. 



Generic diagnosis. — Pupil horizontal. Tongue elliptical, entire and 

 free between the choanae. Fingers free; toes free or very slightly 

 webbed, often fringed; tips of digits not dilated. Outer metatarsals 

 united. Vomerine bone with a backwardly directed process ; quadrato- 

 jugal present, forming a suture %vith the maxillary; omostemum 

 cartilaginous; sternum mth a bony style which is often furcate 

 posteriorly; sacral diapophyses slightly dilated; terminal phalanges 

 simple. 



In Miranda-Ribeiro's original description of Engystomops, spelled 

 Eugystomops, moreirae (1937d, p. 68; type locality, Sorocaba, Sao 

 Paulo), no mention is made of the occurrence of vomerine teeth. A 

 tarsal tubercle and a prominent inguinal gland are present, however. 



