INTERNAL ANATOMY OF THE GAJVIASID^E. 309 



sacculus, and its small third tube entirely coiled away within the sacculus ; hut this, I 

 confess, I am not able to do at present ; it must remain for future investigation. I have 

 little doubt that the third tul)c is tlie homologue of the tube which joins the cornu to 

 the cixmera spermatis in Ltelaps acuta, and it is possible that, under some circumstances, 

 it may emerge from the sacculus and join the camera as in that species. Tlie three 

 chambers of the trefoil arc not usually empty ; they more generally contain very fine 

 granular matter, which might i)ossibly be spermatozoa, but certainly not spermatocysts. 

 It may be possible that in this species, where the male has never yet been found, the 

 sacculus has become modified into an organ wath otlier functions not yet understood ; 

 or it may even be possible that an agamous generation exists, and that the sperm- 

 elements may be preserved in this singular receptacle from the mother Avithin whose 

 body the embryo becomes so liighly developed ; but these are only mere suggestions 

 of possibilities, and I do not in any way put them forward as facts or even 

 probabilities. 



Another totally dilforcnt but very interesting variation of the female genital organs 

 of the Gamasiure from AVinkler's type {G. crass'qics) is found in Scjus togatus, Koch. 

 These parts are delineated in PI. XXXIII. fig. 13. It will be seen how unlike they are 

 to anything yet spoken of; they are a thorough return to the ring form. There is a 

 single azygous central ovary (or), having tlie appearance of a hunch of grapes — all 

 the numerous larger eggs being in oocysts on the exterior, and the germinal part 

 more central. From the ovary proceed two oviducts {od), not one, as iu all the other 

 species we have been considering ; these two oviducts join in an azygous vagina {va), 

 out of which leads an immensely large, flask-shaped spermatheca {spt), which lies 

 between the oviducts, has a narrow mouth and stalk, and powerful muscles attached 

 evidently for the purpose of ejecting the contents. It is generally full of spermatozoa 

 of the curious form shown by fig. 45. These spermatozoa are also found in the 

 oviducts and ovary of the female. 



The principal interest of these organs lies in the fact that, although very unlike Avhat 

 are usually found in the Gamasinae, they closely resemble those of the other subfamily, 

 the Uropodinas; although the general appearance of the creature and the anatomy 

 of the alimentary canal, and most other jjarts, are of the type of the Gamasina?, not 

 of the Uropodinte. This relationship of the female is rendered more interesting from 

 the fact that the male also shows certain affinities to the Uropodimc. 



The Alimentary Canal. 



I have not many variations to record in this system; one or two, however, that I 

 have observed may be of some interest. 



The ventriculus in most species of Gamasidaj is a smallish viscus, having a moderately 

 large and wide csecal prolongation forward in the central line, and often a corresponding 

 one backward. Two large and long paired sac-like cajca spring from its lateral anterior 



