COMPAEATIVE MOEPHOLOGT OF THE GALEODID.E. 353 



small Galeodes, cleared in cedar oil. It shows the base of each haii- suiTounded by a 

 si^ecialized arrangement of cells. Eig. 18 is an enlarged drawing of a few of these, as 

 seen in the cleared specimen, while figs. 14 and 15 rejiresent others in section. 



We find the base of the hair- slung, in a chamber in the solid cuticle, by the hard 

 refractive outermost layer, as above described (p. 333). Its innermost end is attached by 

 elastic (?) fibres to the waU of the chamber. These radiating fibres thus form tos-ether 

 a septum across the chamber, dividing it into an upper and a lower portion. The 

 uj)per portion is kept supplied Avith body-fluid by means of a seimrate channel (fig. 14, c). 

 which I have already mentioned. The hair itself owes its origin to a group of cells 

 which are attached to it only by protoplasmic processes, apjiarently to avoid overcrowding 

 round its small base. One, two, thi-ee, or more nerves run from the base of the hair and 

 swell into large ganglion-cells proximally to the group of the secreting-cells. These two 

 groups together (the secreting- and the ganglion-cells) are protected under a dome-like 

 arrangement of cells (fig. 14, ro) wliich projects into the hollow of the liml), and is closed 

 on all sides except proximally, where the nerves run out to join the nerve-trunks in the 

 limb. Eig. 15 represents a small terminal hair where the protecting roof is not reqrured. 

 The exact innervation of the hair I have not succeeded in making out. I am disposed 

 to think that the nerves terminate in the septum of radiating fibres, and for the follo-vving 

 reasons : — (a) I can find nothing like a nerve running up the axis of the hair ; (6) the 

 radiating fibres would natm-ally be the first to be strained or relaxed — strained on the side 

 towards which the hah- is bent by contact with an object, and relaxed on the opposite side ; 

 ((?) the development of a special channel between the portion above the septum and the 

 cavity of the limb may perhaps imply great physiological activity of this fibrous septum, 

 (6) Other Organs of unknown significance. — At the tips of the pedij^alps and of the 

 pair of legs, which, in the Galeodidse, are gradually losing their claws and becoming 

 feelers [cf. the Pedipalpi), remarkable sensory organs have apparently been discovered 

 simultaneously by Bertkau (20) and Gaubert (32). In each case they occru* on the 

 upper, or dorsal, and slightly on the outer sides of the limbs. They occiu" in con- 

 siderable nimibers, but not in regular arrangement nor in any well-defined areas. 



Gaubert has given a description of these organs, mth a figiu-e showing their distribution 

 on the first leg of Galeodes harharus, and Bertkau has given drawings of the appearance 

 of the organs in section on the j)edipalps of Solpuga flavescens. The observations I have 

 been able to make on these organs are, unfortunately, very far from complete, especially 

 in reference to their soft parts. 

 I have found three kinds : — 



1, (PL XXXII. figs. 11 a, 11 b.) A sort of barrel-like pit in the cuticle, which appears to 

 open externally, and from the base of which a fine pointed process rises, but does not quite 

 reach the external apertiu^e. Passing through the lower boundary of the barrel, the fine 

 process is continued downward into a large bag-like cell, apparently full of fluid. At the 

 base of this cell lies a mass of dense granidar protoplasm with a large nucleus. These bag- 

 like cells are of all lengths, and are supported in a framework of connective tissue, the 

 cells of which often appear to belong to the bags, and indeed I cannot be sure whether 



SECOND SERIES. — ZOOLOGT, VOL. VI. 47 



