500 DE. C. I. FORSYTH MAJOR ON 



In the following year Turner described and figured five distinct distal carpal bones in 

 a Whale. After having mentioned that in 3Iesoplodo)i hldeiis " carpalia 4-1-5 formed a 

 single bone .... which was grooved on its dorsal surface opposite the interval between 

 metacarpals IV and V," he proceeds to describe the carpus of an adult Hijperoodon 

 rostratus. " Tlie distal carpalia are five distinct bones, not so regularly faceted as those in 

 the proximal row, and with a larger proportion of cartilage between them. These bones 

 pass from the radial to the ulnar border in regular order, as C 1 to C 5, and each is 

 associated with the metacarpal bone of its corresponding digit. A similar arrangement 

 exists in both limbs, and the carpus possesses also an elongated pisiform cartilage, which 

 in one is partially ossified " * . 



Von Bardeleben had previously made the following statement : — " Deutliche Anzeichen 

 einer friiheren Trennung in zwei Elemente zeigt das Hamatum bei den Beutelthieren, 

 weniger auff'allend bei den Nagern, sowie bei Ziphiiis {Hyperoodon). In zwei Stiicke 

 getrennt, abcr, auf der einen Seite wenigstens, schon im Verwaclisen begrilTen, ist es an 

 dem Skelete eines jungen Baren in Berlin." t- 



To these assertions Baur replied that he had never in any mammalian embryo observed 

 the hamatum to be the outcome of a fusion of two elements, and he adds : — " Wenn es 

 l)ei iilteren Thieren den Anschein hat, als ware eine Tbeilung vorhanden, so ist dies eben 

 etwas secundiires und ist morphologisch nicht verwendlmr " %. In his latest utterances 

 on the subject §, Von Bardeleben mentions (mly the separation of the hamatum in 

 " Ziphius {llyperoodon)^'' thus tacitly withdrawing the statements regarding other 

 Mammalia, made at the meeting of the Jenaische Gesellschaft of May 15, 1885, above 

 quoted, as %\ell as in a subsequent meeting of October 30 ||. 



The manus of the Jena specimen of Ziphins cavirostris, to wliich Von Bardeleben 

 refers, has been described and figured by Kiikenthal. It contains altogether three distal 

 carpalia : the one resting on Metac. IV and V shows on its dorsal surface a delicate 

 furrow, " eine zarte Furche als Andeutung einer friiheren Trennung zweier Carpalia" ^. 

 This is what Von Bardeleben, in Ids " Referat," calls having found in Zipliim " eine 

 natiirliclie Zerlegung des ' Hamatum ' in das Carpale IV und Carpale V " **, and further 

 on : " Dass Ref. im Mai 1885 die primitive (vielleicht secundare — jedenfalls dem 

 Verhalten bei Urodelen entsprechende) Trennung des 'Hamatum' oder Carpale 4-f-5 

 (Gegenbaur) in Carpale I und Carpale 5 Ijei Ziphius cavirostris autfand (an der Hand des 

 Jenaer Exemplars)." ft. 



In his subsequently-published researches, Kiikenthal describes fresh facts and sums 

 up those previously recorded %%. In embryos of Beluga and Monodon there sometimes 



* Journ. Anat. Physiol, xii. pp. ISO, 183 (188(3). 



t Jeuaische Zeitschr. f. Naturw. xix. (xii.), Suppl. ii. p. 87, Sitzung urn l-j. ilai 188.5. 

 J Zool. Auz. 1885, p. 487. 



§ Proc. Zool. Soc. London, 1894, p. 375 ; " Hand und Fuss," Verh. d. .Inat. Ges. viii. pp. 2()3, 301 (1894). 

 ;i Jen. Zeitschr. xix. (xii.), Suppl. iii. Sep.-Abdr. p. 78 (1885). 



•U Denksohr. d. nied.-naturw. (tes. zu Jena, iii. pp. 38, 4G, pi. iii. tig. 18 (1889). See also E. Rosenberg, op. cit. 

 p. 2, footnote 4 ; Kiikenthal, in Morph. Jahrb. xiv. p. 50 (1893). 



** Op. cit. p. 203. tt 0/:<. C't. p. 301. 



Xt Denkschr. med.-naturw. Ges. Jena, iii. pp. 208-280 (1893). 



