AN ACCOUNT OF SOME HELMINTHES COLLECTED IN THE SUDAN 
197 
genital pore. The gut branches fall short of the anterior limit of the testes hy almost 
a millimetre. 
As before mentioned, the genital system discharges into the spacious ventral pouch, which 
in this species has the configuration found in G. gregarius. The genital pore lies at the 
bottom of a distinct depression, the sides of which are covered with papillae. The posterior 
edge of this depression protrudes so as partly to shut off the inner portion of the ventral pouch. 
The genital apparatus does not appear to differ in any marked respect from that of G. gregarius. 
The vas deferens discharges upon a small genital protuberance. The two testes lie on either 
side and overlap the dome of the posterior sucker. They measure 9 mm. vertically, I'l mm. 
transversely. The disposition of the female genitalia is illustrated in the accompanying Fig. 46. 
It will be noted that the ovarium and shell gland together with the excretory vesicle lie well to 
the dorsal side of the animal and are not wedged in between the dome of the ventral sucker 
and the posterior wall of the ventral pouch. The excretory vesicle is somewhat triangular in 
median section, and its discharging canal tilts a little forwards to meet Laurer’s canal before 
piercing the dorsal integument. In G. gregarius the excretory vessel has a more oval shape 
and its duct passes backwards to open some distance behind the aperture of Laurer’s canal 
(Pig. 45). 
Family, Fasciolid^ 
Genus, Balfouria, nov. 
Balfouria vionogama, n. sp. Plates XXI. and XXII. 
Marabou Stoi’k. Leptoj^tilus crumeniferus, Guv. Intestinal mucosa (encysted). Taufikia. 
In the course of dissection of a Marabou stork. Dr. Wenyon met with an exceedingly 
curious condition of tumour formation upon the peritoneal surface of the intestines. These 
tumours (Plate XXI., fig. 1, to right) are tense and hemispherical, with a smooth glistening surface, 
and measure about half an inch in diameter. They do not invade the mucous membrane of the 
gut to affect it pathologically, but there opens into each from that aspect a tiny pore (Plate XXI., 
fig. 2). When cut into (Plate XXI., fig. 1, on left) each tumour is found to contain quite 
constantly a single pair of sexually mature flukes lying with their ventra in close apposition. 
Such a mode of life is highly exceptional in the DistomidcB, particularly as each worm 
is hermaphrodite and both sets of genital organs are functionally active. Brandes describes a 
somewhat similar condition of encystment for Distomum turgidimi in the intestine of the frog, 
but the number of worms in a tumour varied and each worm was enclosed in a distinct capsule. 
Distomum okenii occurs encysted in the gills of Brama raji, but the parasite is unisexual, 
each cyst containing a single paired male and female worm. The form and structure of the 
parasite in the Marabou stork are so highly characteristic that it deserves, apart from its 
curious mode of existence, to be grouped apart; and I accordingly propose to create a new 
genus, Balfouria, for its reception, whilst mojiogama would probably be an acceptably 
descriptive name for the type species. 
B. monogavia measures about 9 mm. in length, and the body may be recognised as 
divisible into three parts; a pyramidal anterior portion having the mouth at its apex, a rotund 
pea-like central portion filled with ova, and a posterior thumb-like “ appendix ” that is ventrally 
bent and contains the genital and excretory organs (Plate XXII., fig. 1). The ventral sucker 
lies at the junction of second and third portions. Save for the appended third portion, the 
shape of the body recalls that of a Gastrodiscus. 
The anterior half of the body is armed with rows of very stout chitinous spines that 
decrease in size from before backwards, those around the mouth being exceedingly strong and 
arranged after a very definite plan (Plate XXII., fig. 2). 
