CUCULLANIDAE 231 



Fam. 5. CUCULLANIDAE {s.s.) Barreto, 1916. 



Lips large. Mouth a dorso-ventral slit. Oesophagus in 

 two portions, but without a distinct glandular portion, and 

 usually dilated anteriorly to form a large, muscular " false 

 buccal cavity," without a chitinoid lining. Spicules equal. 

 An accessory piece usually present. A preanal sucker-like 

 organ typically present in the male. Vulva in middle region 

 of body, or posterior. Vagina runs forward from vulva. 



1. Cucullanus Miiller, 1777. 



Syn. Pleurorinchus Nau, 1787; Pleurorhynchus Rudolphi, 

 1801 ; Dacnitis Dujardin, 1845 ; Stelmius Dujardin, 1845 ; 

 Dichelyne Jagerskiold, 1902; Bulbodacnitis Lane, 1916; 

 Serradacnitis Lane, 1916. 



Anterior end usually bent dorsaUy. Lips with denticulate 

 border. Preanal " sucker " of male small and fusiform. 

 Caudal alae rudimentary or absent. Vulva very prominent, 

 in posterior half of body. 



Hab. Alimentary canal of Fishes and Turtles. 

 Genotype : C. cirratus Miiller, 1777. 



Miiller, 1777, ZooJogiae Danicae Prodromus, i, 4; Railliet & 

 Hem-y, 1915, Bull. Soc. Path, exot., Paris, viii, 446 ; Barreto, 



1922, Mem. Inst. Osivaldo Cruz, Rio de Janeiro, xiv, 69 ; Baylis, 



1923, Parasitol., xv, 33 ; 1923, Ann. Mag. Nat. Hist., xii, 233. 



The genus Serradacnitis Lane differs only from Cucullanus 

 in possessing " cuticular serration distinct from and in addition 

 to cuticular striation." Bulbodacnitis Lane is defined as 

 " possessing a node or tubercle on the dorsal aspect of the 

 head " (Lane, 1916, Ind. Journ. Med. Res., iv, 97). We do 

 not consider that the characters mentioned warrant the 

 retention of these genera. 



2. Dacnitoides Ward & Magath, 1917. 



Differs from Cucullanus in having no accessory piece, in 

 the form of the uterus and in the presence of an intestinal 

 caecum. The posterior uterine branch ends blindly (as in 

 the family CamaUanidae) and is without an ovary. 



Hab. Intestine of fresh-water Fishes. 



Genotype : D. cotylophora Ward & Magath, 1917. 



Ward & Magath, 1917 (1916), Journ. Parasitol, iii, 60; 

 Baylis, 1923, Parasitol., xv, 31. 



3. Seuratum Hall, 1916. 



8501. Ophiostomum Creplin, 1839, of Seurat, 1915. 



Mouth with two lips. No oesophageal bulb. Tail of male 

 with narrow alae. Spicules short, equal or subequal. An 



