236 DIOCTOPHYMOIDEA 



Mouth hexagonal, surrounded by one, two or three circles 

 each of six papillae. Oesophagus relatively long, simple or 

 club-shaped. Tail of male furnished with a " bursa," without 

 rays. A single long spicule present. Anus of female terminal. 

 Female genital tube single. Eggs barrel-shaped, with thick, 

 pitted, albuminous coating and not containing embryos when 

 laid. Adults parasitic in Mammals and Birds. Larval forms 

 in an intermediate host. 



Fam. 1. DIOCTOPHYMIDAE Railliet, 1916. 

 Syn. Eustrongylidae Leiper, 1908 ; Jagerskiold, 1909. 

 With the characters of the Order. 



1. Dioctophyme Collet-Meygret, 1802. 



Syn. Eustrongylus Diesing, 1851. 



Extremely large forms (up to 1 m. in length in female). 

 Male little more than one-third of the length of the female. 

 Cuticle relatively thin and transparent. Body blood-red 

 during life. Each lateral line with a row of papillae, which 

 become closer together at the extremities. Body truncate 

 posteriorly, somewhat tapering anteriorly. Mouth sur- 

 rounded by two circles each of six papillae, the anterior 

 small, the posterior large. The transversely-elongated, oval 

 " bursa " is indented on its anterior margin, without rays 

 proper, but supported by a triradiate structure in the centre of 

 which is the cloacal opening. The entire margin of the 

 '' bursa " is bordered by papillae. Spicule bristle-like. Tail 

 of female bears a number of small papillae near the anus, which 

 is crescentic. Vulva towards the anterior end of the body, 

 but postoesophageal. 



Hab. Adult : kidney and peritoneal cavity of Mammals. 

 Larva : in musculature of fresh-water Fishes. 

 Genotype : D. [Asccms] renalis (Goeze, 1782). 



Goeze, 1782, Naturg. d. Eingeweidewurmer, 39, 73 ; Collet- 

 Meygret, 1802, Journ. Phys., etc., Paris, Ixv, 458 ; Stiles, 1901, 

 Zool. Jahrb., Syst., xv, 167 ; Leuckart, 1868, Die Menschlichen 

 Parasiten, Leipzig and Heidelberg, ii, 353 ; Dujardin, 1845, 

 Hist. nat. des Helminthes, Paris, 113; Schneider, 1866, Mono- 

 graphie der Nematoden, Berlin, 49. 



2. Eustrongylides Jagerskiold, 1909. 



Forms of medium or rather large size. Body cylindrical, or 

 sometimes swollen in its middle. Cuticle coarsely striated, at 

 least towards the extremities, but without spines. Mouth 

 surrounded by twelve or eighteen papillae, arranged in two 

 or three circles. Vulva close to anus. 



