^BuTDal 



Kmbe' o' Ki|.erii»iint 

 iDortginal account 



8qil. 1 (Table I. p. 36) 



" 9 

 „ 11 

 „ 7 



60 (Tslile VII. p. 67) 



10 (Table I. p. 36) 



4;i iTable IV. p. 50) 

 " K, ITabh. VII. p. 59) 

 " M (Table VI, p. 56) 

 , 70 (Table IX. p. 65) 





: (Tiiblt VJ. p. 55) 



Restatement of earliest Experiments^ carried 02tt before 1902, and descr 



Form of Mating 

 i Hoary x Glabrous sap-colour {CRH) 



[ 



cream {RHK) 



Bap-colour ICRH)] x Glabrous cream (RHK) 



J Hoary x Glabrous white (CK) 



ditto 



[i Hoary X Glabrous sap-colour {CRH)]< 



[..><,. .. „ ]x 



![ I) X II II ,1 ]x I. } X ditto when i'^ = /f 



][ „ X „ ,, „ ] X Glabrous cream (i?H7i.')} x ditto 



U ,, X „ „ „ ]x Glabrous white (CA') } x ditto (?CiC or CHiT) 



when Fo=H 



1[A Hoary x(ilabrous sap-colour (C/fH)]x Glabrous cream (iJHA')J xditto when F;=S 

 [" I, ' X ,1 ,, ,, ]x ditto ixditto ". 



![ I. X , ]x 



([ ,. X „ ,1 „ ]xself 



X ditto 

 X self 

 X self 



J[A Hoary < Glabrous sap-colour {CIlIl)] x self 



} X self when Fj = iH 



60 (Table VII. p. 5G) [4 Hoary -, Glabrous sap-colour (CRU)] :; Glabrous white (C/f) 



51 (Table V. p. 62) 



611 (Table VIII. p. 62) 



•Wr, (Tabic VI. p. 55) Case 3 



[J Hoary x Glabrous sap-colour {CRH)] x 



[ ., X , ]x 



|[ ., X „ „ „ ]x 



} X self when F^ — H 



{[ 



]x ditto 



Case 1 {[i Hoary x Glabrous sap-colour {CRH)} x self 



Case 2 f[ ,, x „ „ i» J*' " 



Case 4 ![ ,, x „ „ n ]** n 



Case 5 {[ „ X „ ,, II ]x .1 



mittee of the Royal Socif.ty. 



\H according to the constitution 

 of the white employed in the 

 hnal mating 



163 

 683 

 637 

 66 



1 KH : -2 iH ■ IG 

 = 1 ; 3 



If, as there is no reason to doubt, all tlie plants employed were 

 homozygous, both the h hoary and the glabrous plants must 

 be due to experimental error. (Hee note 2, p. 148.) 



The ^ hoary condition would uo doubt have been developed 

 if the plants had been allowed to mature. 



The expected equality of hoai-y and glabrous was obtained. 

 As reRards the three J hoary plants see comment on Experi- 

 ments 7 and 11. 



In accord with expectation. 



Commercial glabrous whites have been found sometimes to be a 

 mixture of CK and ( 'HA'. If the white employed in the final 

 mating was CHK in constitution as may quite possibly have 

 been the case— it was not descended from the 'm individual 

 previously used— the result accords well with expectation. 

 We have in fact iudircct proof that such must have been the 

 case, for only on this view should the result which was 

 recorded in a mating; where a fully hoary type was sub- 

 stituted for the half-hoary have been obtained. {See Experi- 

 ment 71 in the original account.) 



in accord with expectation. 



Had the 58 plants recorded as glabrous been allowed to matm'e 

 there is little doubt that the expectation § quarter-hoary 

 and I glabrous wnuld have been recorded. (See similar case 

 of Experiment 20.) The /■'- plant from which F3 was derived 

 was recorded as being distinctly intermediate between the 

 4 hoary and the glabrous condition, i.e. quarter-hoary, and 

 hence would certainly give rise to some quarter-hoary 

 offspring. We may regard the total 58 as representing 

 glabrous and ^ hoary summed together, the ratio of the 

 half-hoary to the sum of tliese two being almost exact. 



In this case a few plants were grown to the flowering stage, but 

 the majority were discarded as seedhnge, with the result that 

 probably ^H as well as ^FI were recorded as glabrous. The 

 proportion of fully hoary to the other three grades taken 

 together is not far out. 



If we bear in mind (1) that here as in the precedingexperiments 

 all the J hoary and a certain number of the J hoary will 

 undoubtedly have been classed as glabrous, and {'2) that the 

 appearance of the form containing the combination JJK is 

 not yet known, and even if distinguishable on inspection 

 from the fully hoary form is likely lo have been classed with 

 that type, we may take these results, so far as they go, as 

 being as near expectation as could be expected under the 

 circumstances. For whereas we have recorded a total of 

 1591H, 107 hH, 98G(; the result of classifying the JJK form 

 with if and the ^7/ with G should give 15it4H, 210 hH, 880(>. 

 The excess of G is no doubt made up of the missing Ul 

 plants which were wrongly classiHed. 



The deviation from a precise result is not greater than one 

 might well expect. 



See remark above. 



In accord witli expectatiou. 



It must be borne in mind that at the earlier date the factor relations underlying these results, which only the fuller knowledge derived from later experiments has made clear, was still matter for investigatiou. 

 uciigh individuals of this constitution no doubt occur we do not yet know whether they are intermediate in appearance between the 4 hoary and the fully hoary ( = .^ hoary), or whether they are iudistinguishable in appearance from one or other of 

 se classes. For the present therefore we can only indicate this form by the distinctive combination of factors— </</ A'. 



