I{. IIak; 'I'lKiMAs 255 



Tlic luiiabur of pluinugu iuuUitiun« in the uuilu.s of Series I recorded 

 in this paper are six : throat, breast, flank, thigh tuft, primary and 

 central rectrix, the five first transmitted through four generations, the 

 sixth appearing first in F... Their description in (letail will be found in 

 the Appendix. 



To fully appreciate the importance of these fixed mutations in four 

 of these areas of the males of Series 1, we must refer to the description 

 of the males of the two species crossed. We find in both of them that 

 all the inferior parts are unpatterned, yet we get a fixed patterned 

 condition of the inferior parts in F^ " BBA," F, " BBBA " and 

 Ft " BBBA " inter se with a certain segregation showing that the un- 

 patterned condition has become recessive. 



Taking the four areas of the inferior parts— throat, breast, flank 

 thigh tuft separately, and making the reckoning from three generations, 

 we find the segregation in them is as follows, F„ "BBA," F., "BBBA " 

 and Fi " BBBA " inter se, eighteen males. 



Males: 1<'.,''HISA,'' 5\ F^-'IilWA" 8; Fi'-BltlSA" 

 Source of mutatiou, male Swiulioe iutL'rscapul!ir> 



Source of mutation, female Swinhoe breast and 

 flank 



Totals ~. 55 11~" 



The fifth mutation in the mules of Series 1 was the central rectrices 

 which in F^ " BA," F„ "BBA" is found heavily patterned, while in 

 F^ " BBBA " and F^ " BBBA " inter se Mendelian segregation occurs, 

 one individual having the absolute male Swinhoe type central rectrix 

 which is unpatterned white with white rachis. 



In this fifth area a patterned takes the place of an unpatterned 

 condition, as in the first four mutations. 



In the sixth area, the primaries, two changes occur — a marked 

 reduction in the number of bands, and the breaking up of these into 

 dots and dashes, with proportionate increase of the white area; here 

 the tendency is for pattern to disappear rather than t(j increase. 



It should be emphatically stated that in Series 1 male mutations 

 throat, breast, and flank, the pattern does not in the remotest degree 

 resemble the transitional plumage patterns found in two of those areas 

 on the young male Silver: therefore arrested development cannot 

 explain the phenomena. The origin is clearly traced to the male 



