﻿330 SMITHSONIAN MISCELLANEOUS COLLECTIONS [VOL. 47 



ably more lenticular than any of the small wingless seeds from the 

 Coal Measures placed by authors in Carpolithes. As between 

 Lagenostoma and Rhabdocarpos, the two types of seeds reported to 

 have been definitely correlated with Cycadofilic fronds, it is evidently 

 with the former that the closer relations probably exist, although it 

 is difficult to determine satisfactorily the relations between seeds 

 known chiefly by their internal structure and others known only 

 by the external characters of the compressed specimens. The most 

 prominent differences between Lagenostoma, as the latter is de- 

 scribed by Williamson 1 or Oliver and Scott, 2 and Wardia are the 

 presence of the wing, the thin and lenticular cross-section, and the 

 absence of all traces of radial chambering near the apex, unless the 

 faint costation is to be interpreted as due to the features of the 

 pollenic chamber. Wardia has probably two distinct tests ; Lagenos- 

 toma physoides, which has a fibrous layer, perhaps approaches more 

 closely than do L. ovoides or L. Lomaxi, the latter of which is pro- 

 duced in a costate and deeply lobed cupule. Certain points of re- 

 semblance between the two genera include a degree of similarity in 

 size and general form of the seeds, the development of the latter 

 on pedicles, and, to some extent, a common Spenopteroid habit of 

 the sterile fronds. Incidentally the punctation on the rachis and the 

 pedicels of Wardia may be homologous with the glandiferous stalks 

 on the pedicel and the cupule of Lagenostoma. On the other hand 

 the reduced pinnules of Aneimites fertilis are in general form very 

 suggestive of those of the normal Sphenopteris elegans, with which 

 Heterangium, another Pteridospermic type, is correlated, while the 

 narrow, ventrally sulcate, dorsally round divisions of the Aneimites 

 rachis suggest the V-shaped single vascular axis of a Heterangium 

 petiole. 



On account of certain similarities between the sterile fronds of 

 Aneimites and those of the elegans and Hoeninghausii groups of 

 Spenopterids, together with the consideration of the points of resem- 

 blance in the seed production in Lyginopteris, I am disposed to 

 regard Wardia as more closely related to Lagenostoma than to any 

 other Cycadofilic type at present recognized. But, in the absence of 

 knowledge of the internal organization of the rachis of Aneimites, 

 and in view of the important external seed differences, the inclusion 

 of the latter genus within the same family (Lyginodendreae) does 

 not at present seem warranted. However, they may, I believe, with 



1 Trans. Roy. Soc, vol. clxvii, 1877, p. 233, figs. 53-75; vol. clxx, 1880, p. 

 517, figs. 61-63. 



2 Proc. Roy. Soc, vol. lxxi, 1903, p. 477; vol. lxxiii, 1904, p. 4. 



